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GREAT BLUE HERON <br />Predation. Northwestern Crows (Corvus <br />caurinus) and Common Ravens (C. corax) eat <br />unattended eggs (Butler 1989). Predators of <br />nestlings include eagles (Kelsall and Simpson 1979, <br />Koonz 1980, Forbes 1987, Norman et al. 1989), <br />raccoons (Lopinot 1952, Hjertas 1982), bears (Foss <br />1980, Parker 1980), Turkey Vultures (Meitner 1951) <br />and Red - tailed Hawks (Buteo jamaicensis;Simpson <br />1984). Colony -sites abandoned after predators kill <br />adults (Butler 1991) and nestlings (Kelsall and <br />Simpson 1979, Simpson et al. 1987). <br />BREEDING <br />MATE SELECTION <br />Little information. Of 5 marked pairs in 1978, <br />all had new mates the following year (Simpson <br />1984). Herodias and occidentalis form pairs and <br />produce viable offspring with white and blue <br />plumages (Bent 1926, Powell et al. 1989). <br />PHENOLOGY <br />Pair formation. At some northern latitudes, <br />adults gather in flocks on the ground for several <br />days in spring before entering colony -sites <br />( "gathering grounds" sensu Mock 1976). Birds do <br />not occupy gathering grounds in Texas and <br />Michigan, and some roost at colony -site year - <br />round (Cottrille and Cottrille 1958, Mock 1976). <br />Function of gathering ground unclear. <br />Little published information on dates of pair <br />formation. Earliest adults return to colony -sites as <br />follows: California, Dec (Brandman 1976); Texas, <br />Jan (Mock 1976); British Columbia, mid -Jan on the <br />coast, late Mar interior sites (Butler et al. 1986, <br />RWB, Cannings et al. 1987); Pennsylvania, mid - <br />Feb (E. Bruckner pers. comm.); Alberta, late Mar <br />(Vermeer 1969). Herons . in some colonies apt'to <br />depart from colony -sites with little provocation at <br />this time;: may be inhibited from displaying by <br />high winds and low temperatures (Palmer 1962). <br />Courtship to egg laying early Jan to mid -Mar in <br />California (Brandman 1976), mid -Feb to early Apr <br />in British Columbia (RWB). Great White Heron <br />(occidentalis group) breed year -round but most <br />begin between Sep and Feb (Powell 1983). <br />First brood per season. Figure 3. First eggs: late <br />Feb in Oregon (Henny and Bethers 1971) and n. <br />California (Pratt 1970, Pratt and Winkler 1985); <br />early Mar in s. California (Brandman 1976); 3rd <br />week Mar in Idaho (Collazo 1981); 1st week Apr in <br />s. British Columbia (Butler 1989); mid-Apr in Nova <br />Scotia (McAloney 1973); late Apr in Alberta <br />(Vermeer 1969). <br />The Birds of North America, No. 25, 1992 <br />Imo Primaries <br />Molt Body <br />Breeding Young <br />E <br />Mgration 7:11 <br />Figure 3. <br />Annual cycle of <br />breeding, molt, and <br />migration of Great <br />Blue Herons In <br />British Columbia. <br />Coastal populations <br />are non-migratory. <br />Thick lines equal <br />peak activity, thin <br />lines off peak. <br />A. Poole, P. Stettenheim, and F. Gill, Editors <br />Second brood per season. Unclear how many <br />late -laid Butches are first or second attempts. Two <br />pairs renested after fledging first brood, 5 pairs <br />after death of nestlings in California (Brandman <br />1976). Most choose new mates after first attempt <br />fails (Simpson 1984). Insufficient time to raise 2 <br />broods at northern latitudes. <br />NEST SITE <br />Nest as single pairs but mostly in colonies (Fig. <br />4). Mean distance flown from colony to principal <br />feeding sites 2.3 to 6.5 km (Dowd and Flake 1985b, <br />Thompson 1978, Parris 1979, Butler 1991). Two <br />radio - equipped breeding adults travelled 61.8 and <br />103.7 km in 20 to 25 h (Parris 1979). Number of <br />nests in British Columbia (Butler 1991), Oregon <br />(Werschkul et al. 1977, Bayer and McMahon 1981), <br />and Maine (Gibbs et al. 1987, Gibbs 1991) positively <br />related to area of nearby foraging habitat. Location <br />of colony sites best explained by distribution of <br />foraging habitats (Butler 1991, Gibbs 1991). Site <br />selection also predator - driven; like most other <br />herons, this species generally selects nest sites <br />difficult for mammalian predators to reach, e.g., <br />islands, trees in swamps, high branches, etc. <br />Recovery of beaver (Castor.canadensis)populations <br />in ne. U.S. and s. Canada may have benefited this <br />species by providing a patchwork of large and <br />small swamps and wet meadows — additional <br />nesting and foraging areas (e.g., N.Y. State; Andrle <br />and Carroll 1988). <br />