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HomeMy WebLinkAbout01/30/2002 Env Bd PacketCity of Lino Lakes Environmental Board Meeting January 30, 2002 6:30 p.m. AGENDA 1. CALL TO ORDER/WELCOME NEW MEMBERS 2. APPROVAL OF MINUTES 3. APPROVAL OF AGENDA 4. OPEN MIKE 5. ACTION ITEMS A. Election of Officers 6. DISCUSSION ITEMS A. Wildland Urban Interface /Completed Project Report/Peggy Booth, Minnesota Department of Natural Resources B. Peltier Lake /Proposed Boating Restrictions/Heron Nesting Area C. Development Regulations Task Force 7. DEPARTMENT REPORTS A. Forestry B. Solid Waste Recycling/Solid Waste Report July 1, 2001 to December 31, 2001 8. ADJOURN Agenda Item 6A Environmental Board Meeting Date: January 30, 2002 Topic: Ordinance Proposal /Peltier Lake Use Regulation Background: The Peltier Lake Island is classified as the second largest nesting site for the Great Blue Heron. The island is also a nesting area for the Great White Egret. Records of these birds nesting habits in Lino Lakes have been maintained since 1945. Before 1979 there were two other sites in Lino Lakes that the birds used for nesting. One was on the East Side of Rice Lake and the other was in the Lamprey Pass area. From 1979 to 1991, the birds from these two sites appeared to have migrated to the Peltier Lake nesting site. The theory is that the birds moved away from development in the areas. The first record of colonies on Peltier was recorded in 1989. The records indicate the decline in population of the Rice Lake and Lamprey sites, and in increase in the population of the Peltier site. In 1999, there were 625 recorded nests on Peltier Island. At that time, the Peltier colony became the second biggest colony in the state. The Pigs Eye colony was the largest. There are only five or six nesting colonies in the Metropolitan Area. In 1996 there were 1100 recorded nests on the island. In 1998, a storm reduced the number of nests to 500. The birds were nesting at the end of May 2000. After June 14, the birds were gone. A number of issues may have caused the disappearance of the Blue Herons. Last year was a budget wildfire year. The DNR was practicing aerial water pick- ups on Peltier during the nesting time. Highway construction began on Interstate Highway 35W. The Highway project occurred within % mile from the nesting birds. Aerial photographs show about 60 nesting Egrets on April 16, 2000. The birds left sometime after this. The Blue Heron numbers also went down. In 1998, a permit was issued for private water ski slalom coarse. It is not known if the permit for this coarse was issued in 1997, or weather skiing was occurring at that time. The coarse was set up on the North side of the island. The birds nest on the North side of the island. Research indicates the need for at least a 300 -meter nesting and buffer zone for the Great Blue Heron, and that the birds tend to abandon nests with larger disturbances within 500 meters Butler. 1992. A task force of stakeholders met on January 23, 2002 to discuss action that might be taken to protect the Great Blue Heron Rookery on Peltier Island. One of the suggestions is to immediately communicate with the DNR fire fighters, and the Minnesota Department of Transportation concerning Heron sensitivity issues. Discussions with the skiing group will also occur. It is thought that if a "handshake agreement" can be reached with the skiers to not use the area during critical times, then observations can be made of cause and effect. The other issue that was discussed was lake -use regulation. Regulating the use of Lakes in Minnesota is delegated to local govemments. Regulation is achieved through local ordinance establishment. There are two local govemments involved in the issue of regulating use on Peltier Lake. They are the Cities of Lino Lakes and Centerville. The two cities would need to pass joint powers agreement and ordinance. The proposed ordinance would include a no -wake zone in the area to the North of the Southem section lines of Sections 10 and 11. This would be a line east and west starting at the southem boundary of Rehbeins Peltier View Subdivision of Lino Lakes. Biologists at the task force meeting have indicated that we have one, maybe two more chances to save the Heron Rookery. Analysis: Without some action taken very soon, the City of Lino Lakes is in danger of loosing a unique natural resource feature. The Herons start nesting in April, any regulation before this time would ensure the buffer /use needed to encourage the retum of the Colony. Another side point of high -power boat prop use in the area North of the island is that of water quality impact. Normal water depth in this area is registered as 4.5 feet. During low -lake elevation periods, this can be less than 3 feet. Aerial photography shows channeling and tum- around curl in the ski coarse. Sediments are very phosphorous rich in our lakes. The effect of the prop churning of these sediments is that of pumping phosphorous into the water column. Phosphorous is the limiting factor to plant growth in our lakes. Although unverified by phosphorus sampling data, observations made indicated an unusual increase in algae blooms in the lake after skiing events. Options: Proceed with a joint - powers ordinance regulating use of Peltier Lake in the island area. Recommendation: Proceed and process this regulation language as soon as possible. Attachments: 1. Butler Technical Report 2. DNR Water Surface Use Summary 3. Lablanc Report Ardea herodias FRENCH: Grand Hiron SPANISH: Garza morena, Garza Manta granda, Gatlinaza Great Blue Heron The Great Blue Heron is one of the most widespread and adaptable wading birds in North America. Up to seven subspecies have been recognized by past researchers, based on differences in size and plumage color, but a single subspecies (herodias) probably suffices for most of the continent, except for Florida's Great White Heron (occidentalis), the subspecies most distinctive in color (entirely white). Occidentalis interbreeds freely with herodias to produce an intermediate form, Wiirdemann's Heron of the Florida Keys. This account focuses on both of these subspecies: the continental Great Blue Herons (A. h. herodias), sometimes referred to as the herodias (or blue) group, and the Great White Heron, the occidentalis (or white) group. Both have received considerable attention from researchers. This species nests mostly in colonies, usually large ones of several hundred pairs. Such colonies are often located on islands or in wooded swamps, isolated locations that discourage predation by snakes and mammals. Although this species is primarily a fish eater, wading (often belly deep) along the shoreline of oceans, marshes, lakes, and rivers, it also stalks upland fields for rodents, especially in winter. Its well- studied, elaborate courtship displays have correlates on the foraging grounds, where this species can be strongly territorial. ROBERT W. BUTLER The Birds of North America Life Histories for the 21st Century Figure 1. Breeding, nonbreeding, and year -round ranges of A. herodias. The species is rare in winter in the northern parts of its range. Order CICONIIFORMES Family AREIDAE GREAT BLUE HERON Equally at home in coastal (marine) environments and in fresh water habitats, the Great Blue Heron has weathered the impacts of 20th century North Americans quite successfully, although its breeding colonies remain vulnerable to dis- turbance. DISTINGUISHING CHARACTERISTICS Largest heron in North America, about 60 cm tall, 97 to 137 cm long, 2.1 to 2.5 kg mass. Middle toe with small comb (pectinate). Wings long and rounded, bill long and tapered, tail short. In flight, folds neck in S -shape and extends legs along the body axis; deep, slow wingbeats. Herodias (blue) group: legs and neck long. Long body and occipital plumes on adults. Upperparts gray, fore -neck streaked with white, black, and rust- brown. Bill yellowish. Legs brownish or greenish. Occidentalis (white) group resembles herodias except wholly white (some individuals have a few dark feathers). The Great Blue Heron resembles the Grey Heron of Europe and Africa, found occasionally in the West Indies (American Ornithologists' Union 1983), except that Great Blues have gray feathers with violaceous tinge on back and sides of neck, chestnut feathers on thighs. Adult Grey Heron slightly smaller (90-98 cm long), with pale &ray neck and white feathers on thighs (Hancock and Kushlan 1984). DISTRIBUTION 3U CHECK -LIST REGION Breeding range. Widespread (Fig. 1). The Great ue Heron (Herodias group) nests as single pairs d small colonies along coasts of se. Alaska (61°N; . Sigman pers. comm.) and n. British Columbia WB). Mostly in colonies on south coast Butler 1989) and mountain valleys of British Columbia, in central Canadian Prairies (Vermeer 1969,1970, 1973, Vermeer and Anweiler 1970), s. Ontario (Gray et al. 1980, Dunn et al. 1985), s. Quebec (DesGranges et al. 1979), and the Canadian Maritime provinces (McAloney 1973, Quinney 1982) except Newfoundland (Montevecchi and Tuck 1987), south to Florida, Texas, Baja California, and Central America at least to Belize and Guatemala. Mostly in colonies and along the Pacific and Caribbean coast of Mexico to Guerrero (AOU 1983). Breeding status unknown in central Mexico. Occidentalis group (Great White Heron) restricted to coastal habitats of s. Florida (including the Keys), Cuba, the Isle of Pines, St. Thomas, and The Birds of North America, No. 25, 1992 Anegada (Hancock and Kushlan 1984). Formerly in Jamaica (Spendelow and Patton 1988). Winter range. Pacific coast south of 61 °N (Williamson et a1.1965) through Central America, and mostly south of Canada and midwest U.S. states (Fig. 1). Includes the islands and coast of the Caribbean south to Colombia (Hancock and Kushlan 1984). RANGE OUTSIDE AOU CHECK -LIST REGION Great Blue Heron breeds on the Galapagos Islands (Harris 1973), Great White Heron on islands near Venezuela (Hancock and Kushlan 1984). Former casual in winter in Venezuela and Colombia (Byrd 1978, Hancock and Kushlan 1984); stragglers recorded on Hawaiian Islands (Berger 1972). FOSSIL HISTORY Large herons referable to the living genus Ardea have been in existence since the middle Miocene and probably before, although the phylogenetic relationships of fossil species to living species is still not clear. Earliest record for Ardea is an undescribed species from the early Barstovian North American Land Mammal Age (NALMA: 14 million yr before present) from Observation Quarry, Dawes Co., NB, about the size of the Great Egret (Casmerodias albus; Becker 1986). Next record for Ardea sp. from the late Clarendonian NALMA (10 mybp) from Love Bone Bed, Alachua Co., FL, the size of A. h. occidentalis (Becker 1985b). Ardea polkensis Brodkorb (1955:17), the only correctly named fossil species within the geographic range of the living species (Olson 1985: 165-168), from the late Hemphillian NALMA (5 mybp; Bone Valley, Polk Co., FL); was smaller thanA. herod ias. A late Blancan NALMA (2 mybp) Ardea sp. record, recently made by Emslie (1992) from the Macasphalt Shell Pit, Sarasota Co., FL, was slightly smaller than female herodias and equal in size to the White- necked Heron, A. cocoi. Fossils of A. herodias recorded from many Pleistocene (1.8 mybp) and prehistoric sites within the U.S. (see Becker 1982: 449,1984: 203, 1985a: 38, also Brodkorb 1963: 284, Guthrie 1992: 321, Parmelee 1977: 200, 1985: 176, Howard 1969), Mexico (Hamblin and Rea 1985), and St. Croix in the West Indies (Wetmore 1937). SYSTEMATICS The Great Blue Heron has two close relatives, the South American Cocoi Heron (A. cocoi) and the Old World Grey Heron. These three taxa are highly similar morphologically (Bock 1956), behaviorally The American Ornithologists' Union (Curry- Lindahl 1971), and genetically (Sheldon 1987), suggesting that they constitute a single species. Because they do not overlap in distribution, however, these taxa are usually considered to be semispecies constituting a superspecies (e.g., Hancock and Elliot 1978, Payne 1979). Such a classification permits each of the semispecies to be divided into subspecies. Hancock and Elliot (1978) divided Great Blue Heron into seven subspecies, wardii (se. North America),cognata (GalapagosIs.), treganzai (middle North America), hyperonca (w. U.S.), sanctilucae (s. California), and occidentalis (Florida and West Indies). They also suggested that two more subspecies might usefully be described for Central American and Carribean forms. In general, most researchers view occidentalis (Great White Heron) as a localized color morph, but Curry- Lindahl (1971) believed it differed from typical Great Blue Herons in its behavior, possibly enough to be a distinct species. Payne (1979) recognized four subspecies, fannini, herodias, cognata, and occidentalis. He noted that while southern birds are larger than northern birds, the eastern variation in size is clinal. Thus, a single subspecies (herodias) suffices for most of continental North America, with the exception of the northwest (fannini) and Florida ( occidentalis). This account treats occidentalis as a color morph /subspecies of A. herodias. MIGRATION NATURE OF MIGRATION IN THE SPECIES Migrates alone or in groups of 3 to 12, occasionally up to 100, day and night (Palmer 1962). Some wander northward in summer to arctic Alaska, s. Yukon, s. Keewatin, n. Manitoba, n. Ontario and n. Quebec (Brock 1959, Godfrey 1986). Southward migration from northern localities from mid -Sep to late Oct. General movement away from northern edge of breeding range for winter (Henny 1972), but some recorded on Christmas Bird Counts in Canada each year. Spring migrants return in early Feb to Illinois, Wisconsin, and central Minnesota (Palmer 1962); mid -Mar to Vermont (Laughlin and Kibbe 1985) and British Columbia Butler et al. 1986); late Mar to Kentucky (Mengel 1965), Iowa (Dinsmore et al. 1984), and Oklahoma (Sutton 1967); early Apr to Nova Scotia (Tufts 1 %1), Ontario (Devitt 1967), and Alberta (Vermeer 1969). Also arrive Nova Scotia by Mar, Canadian Prairies and Maritimes in Apr and early May (Palmer 1962). More infor- mation on arrival and departure dates needed to establish migration chronology. ROBERT W. BUTLER TIMING AND ROUTES OF MIGRATION Little information available. Most probably winter along ice -free coastlines and watercourses. Recoveries of U.S. and Canadian herons banded as nestlings east of the Rockies and recovered before their first birthday suggest many winter along Caribbean shores (Byrd 1978). Christmas Bird Counts (CBC) show large numbers in se. U.S. (see Fig. 5). Pacific coast populations appear nonmigratory (Byrd 1978, Gill and Mewaldt 1979) but some post- breeding dispersal occurs (Pratt 1970, Butler 1991). Large concentrations in Puget Sound, WA, and Strait of Georgia, BC. MIGRATORY BEHAVIOR No information. CONTROL AND PHYSIOLOGY OF MIGRATION No information: HABITAT BREEDING RANGE Widespread and remarkably adaptable. Feeds mostly in slow moving or calm freshwater, also along seacoasts. Occasionally in surf and fields. Nests in trees, bushes, on ground and artificial structures (see Breeding: nest site), usually near water; prefers vegetation on islands or in swamps, probably to avoid ground predators. Breeding at elevations up to 1,100 m in British Columbia (Campbell et al. 1990), 610 min Vermont (Laughlin and Kibbe 1985), and 1,500 min Panama (Hancock and Kushlan 1984). Along east coast of U.S., avoids nesting in marine habitats in New Hampshire and New York (Spendelow and Patton 1988), favoring inland sites; farther north and south, nests near fresh and salt water. Great White Heron found almost exclusively in shallow -water marine habitats, particularly tidal grass flats. SPRING AND FALL MIGRATION No information but probably similar to breeding season. WINTER RANGE Little information. In Oklahoma, avoids farm ponds with little emergent vegetation in favor of natural wetlands and riverbanks (Heitmeyer 1986). In British Columbia, flies to estuaries (adult females) and nearby grasslands (juveniles) in autumn and winter when high tides and declining fish populations make foraging unprofitable on beaches; some adult males spend fall and winter on territories along river banks (Butler 1991). Also The Academy of Natural Sciences of Philadelphia GREAT BLUE HERON forage occasionally in dry fields. Along east coast of U.S., favors coastal marine habitats, especially salt marshes. FOOD HABITS FEEDING Main foods taken. Mostly fish but also amphibians, invertebrates, reptiles, mammals, and birds (Palmer 1962, Kushlan 1978, Verbeek and Butler 1989). Microhabitat for foraging. Different foraging habitats best explained by individual ability. In British Columbia, juveniles forage in grasslands, adult females in estuarine marshes and intertidal beaches, adult males along riverbanks (Butler 1991). Microhabitats of yearlings poorly kn onn; include estuaries and beaches, p l visits to colony -sites (RWB). Food capture and consumption. Forage singly and with conspecifics; behavior flexible, diet varied. Hunt most often by slowly wading or standing in wait of prey in shallow water ( Kushlan 1976, 1978, Willard 1977, Hom 1983); also hunt from floating objects (Godin 1977) and even -while floating;:SJensen 1932). Wade more slowly and stop for longer periods than other herons (Willard 1977). Often forage in flocks with other Great Blue Herons (Krebs 1974), other ciconiiformes (Willard 1977, Kushlan 1978), and Double- crested Cormor- ants (Phalacrocorax auritus; RWB). Forage -night and day on beaches (Black and Collopy 1983) and wharves (RWB); up to' one :third-- of total photoreceptors in ciconiiform eyes :.are: rods, presumably an adaptation fornight vision (Lish 1982). Prey located by sight (Krebs and Partridge 1973) and caught by rapid forward thrust of neck and head; then held between Mandibles. Most prey swallowed whole. Small mammals sometimes wetted before swallowing (Peifer 1979, but see Bayer 1981a). Armoured fish taken ashore, then speared and shaken with beak to break or relax the spines (Forbes 1982, but see Bayer 1985a). DIET Major food items. Wide array of animals induding fish, insects, mammals, amphibians, and crustaceans (Willard 1977, Kushlan 1978, Peifer 1979). Voles 24% to 40% of diet of nestlings in Idaho (Collazo 1979); also important to juvenile survival in British Columbia in winter (Butler 1991). Quantitative analysis. Data scant, mostly from breeding season. Predominantly fish (Parker 1980, Quinney and Smith 1979, Parker 1980, Horn 1983, The Birds of North America, No. 25, 1992 Butler 1991). Occasionally chokes to deathonlarge prey items (Wolf and Jones 1989). NUTRITION AND ENERGETICS Estimated mean (± SE) intake of metabolized energy per day by individual herons feeding on small fish during 4 breeding stages was:' egg- laying 1,163 kJ (± 555), incubation 1,197 kJ (± 194), small chicks 4,264 kJ (± 764) and large chicks 1,598 kJ (±`151; Butler 1991). DRINKING, PELLET CASTING, AND DEFECATION Water probably from diet. Casts pellets of mammal hair. Bones digested. Territorial herons depart from foraging sites to defecate more often than nonterritorial herons (Bayer 1980). FOOD SELECTION AND STORAGE Few data; fish about 5 to 30 art long, occasionally longer (Willard 1977). SOUNDS VOCALIZATIONS Mostly silent except at breeding colonies and when disturbed on foraging grounds. Geographic variation unknown. Calls show much variability and intergrade (Bayer 1984a). Vocal array. Bayer (1984a) lists 7 calls at breeding colonies: Frawnk call: rapid squawk, kt average duration of 19.7 s, given day when alarmed or when aggressive toward conspecifics; it may account for the local name "Crank" given to this species along the New England coast. Go -go-go call: a series of ducks given at the foraging site and breeding colony and often answered with same. The awk call: a scream lasting an average of 2.3 s, given mostly inbreeding colonies. The goon call resembles the bleat of a calf; uttered at the end of the "full forward" display (see Fig. 2a). Theee call: divided into two segments, uttered day and night mostly while flying. The roh- roh -roh call: a series of squawks uttered spontaneously for about 3.6 s by herons on the feeding grounds; also arrival at nests is often preceded by this call. Perhaps advertises territorial ownership on the foraging ground (Bayer 1984a) and mate recognition at nest (Mock 1976). NONVOCAL SOUNDS Loud bill snap, part of sexual display (see Behavior: sexual). Males snap bills most often when unmated and defending a nest site; also during bachelor stage when displaying toward females (Mock 1979); also once pairs form, but less The American Ornithologists' Union often than at other stages. Females snap bills when approaching bachelor males and after they have formed breeding pairs (Mock 1979). Possibly analogous to territorial song of passerines (Mock 1976). Bill clappering (Hancock and Kushlan 1984), a rapid chattering of the tips of the bill, is very common between paired birds; many other heron species also do it (Hancock and Kushlan 1984). BEHAVIOR LOCOMOTION Walking. Walks erect with long strides and wades in water, often up to belly or nearly so. Toes are spread as the foot is placed on the ground. Flight. Folds neck in S -shape and extends legs along the body axis on long flights. Extends neck and dangles legs when preparing to land, when chasing other herons, when startled (e.g., when pursued by Bald Eagles, Haliaeetus leucocephalus), and at colony -sites during courtship flights. Flies with deep, slow wingbeats at 2.3 to 3.2 beats /s; flight speed 30 to 46 km /h (Palmer 1962). Soars to great heights and glides occasionally. Capable of evasive maneuvers when pursued. SELF - MAINTENANCE Preening, head- scratching, stretching, etc. Powder downs on flanks and pectinated middle toe nail used for scratching are common to herons, including this species. Stands on one foot to scratch head by lifting leg over closed wing, erecting crest feathers, and lowering and twisting head in position where it can be reached by pectinate toe. Preens by sliding bill along feathers. Droops wing slightly when preening neck. Partly opens wing when preening underwing feathers. Shakes head, body, and finally tail in one motion. Cleans bill by wiping on branches or shaking. Sunbathing, thermoregulation, temperature metabolism. Droops and exposes inside of wings on sunny days, perhaps to radiate body heat on warm days and absorb solar radiation on cool days (Kahl 1971, Larkins 1989). 'Avoids strong winds by seeking shelter behind bushes, fences, etc. Leaves nest to drink on hot days (Pratt 1977). Sleeping and roosting. Roosts alone or in loose flocks of over 100 on the ground, in trees, and on man -made objects near feeding grounds during day. Some roosts used repeatedly. Sleeps at night in trees with dense foliage. Tucks bill under wing - coverts while asleep. Daily time budget. On Pacific Coast, activity controlled mostly by tides. Peak feeding activity near low tide (Brandman 1976, Bayer and ROBERT W. BUTLER McMahon 1981); pair formation, courtship, and nest building generally just before or after foraging (Brandman 1976). During pair formation male spends 100% of his nonforaging time at the nest (perhaps to guard against robbing of nest sticks), 50% to 60% during courtship, 100% during incu- bation, and 51 % at nest when chicks are > 4 to 5 wk old (Brandman 1976). Female spends 50 to 60% of her nonforaging time at nest during courtship, 100% brooding chicks < 4 to 5 wk old, and 31% with older chicks (Brandman 1976). Adult nest attentiveness declines from about 83% to 99% in the first week after hatching to near zero at 6 to 7 wk (Dowd and Flake 1985a). Feeding- activity peaks when chicks are 4-wk old; Males spend less time foraging and more time on the nest than females during the day (Brandman 1976); opposite at night. AGONISTIC BEHAVIOR Physical interactions. Individuals erect crest and fly toward conspecifics that approach within about 2 m of nests and nest sites; also jab such intruders with bill during courtship (see Fig. 2c). Thrust bill at herons that approach nest after eggs are laid. Communicative interactions. The rich repertory of courtship displays at the nest are described below (see Behavior: sexual); displays in this section described by Bayer (1984b). The most spectacular display given by herons on the foraging grounds is the "Upright and Spread Wing" display given when two herons approach one another; neck is nearly fully extended and tilted slightly over the back, head and bill are held above horizontal, wrists are drooped or held away from the body exposing the black shoulder patch of the adult. Sometimes wing nearest the opponent is drooped lower than the opposite wing and body plumes are erected. As opponents approach one another, head and bill are raised closer to the vertical and the neck extended farther over the back. These displays usually cause the two herons to move apart but occasionally bring a chase. This display is sometimes directed towards gulls (Larus sp.), Great Egrets (Casmerodius albus), and even people. A less spectacular display on the foraging grounds is the "Vertical" display (Bayer 1984b). A heron performing this display sleeks its plumage, extends the neck forward at about a 45 °, and tilts the head along its axis so its eyes alternately direct upwards. This behavior used when predators and herons fly at high altitudes over the foraging grounds. The "Forward and Full Forward" display entails partly folding the neck with the bill facing the opponent and partly erecting the body plumes. The Academy of Natural Sciences of Philadelphia GREAT BLUE HERON A From this position, an aggressor can thrust the bill at an opponent, although such attacks are not frequent. The Full Forward behavior, however, can result in serious injury and even the death of an opponent (Forbes and McMackin 1984). "Bill duels," in which opponents alternately attempt to C The Birds of North America, No. 25, 1992 B Figure 2. Breeding displays of the Great Blue Heron. (A) Forward display; (B) Stretch; (C) Bill duel. By D. Otte, from Mock (1976). A. Poole, P. Stettenheim, and F. Gill, Editors grab each other's head (Fig. 2c), occur during some Full Forward displays. "Circle Flights" involve a heron taking flight with the neck extended and parallel to or below the axis of the body; used by territorial herons along boundaries and by nonterritorial herons pursuing one another. In the "Arched Neck" display the neck is fully extended above or parallel to the body axis with a down -curve in the distal portion; used by territorial herons after preening, stretching, or long periods of inactivity, during short flights within a territory, while walking and wading, and while flying to a roost or colony site. Also used by herons in pursuit flight, when flying from an approaching heron or predator, or when supplanted by another individual. Displays on the foraging grounds may be analogous to displays at the nest (R. Bayer pers. comm.). Communicative interactions between chicks and their parents need study. SPACING Individual distance. Nonterritorial herons from neighboring colonies overlap foraging grounds (R. Bayer pers. comm.). Individual distance on foraging grounds and at nests maintained by displays and aggressive interactions (Bayer 1984b, Mock 1976). Territoriality. Adults defend territories along rivers, creeks, mudflats, and lografts day and night (Bayer 1978, Butler 1991). Territories maintained by displays, threats, and chases (Bayer 1984b). Territorial herons rest in exposed sites from where they display and launch chases of approaching herons (RWB). Mean length of shoreline of 32 territories in Yaquina estuary, OR 355 m (SD = 168); mean area: 8.4 ha (SD = 5.4); in freshwater marshes (n = 7 territories): 129 m (SD = 28); mean area: 0.6 ha (SD = 0.1; Bayer 1978). The American Ornithologists' Union Number of territories declines through winter; 14% to 20% of territories in Yaquina estuary are defended by juveniles in Oct, but none in winter (Bayer 1978). On the Fraser River Delta, BC, adult females and juveniles feed nonterritorially on beaches, estuarine marshes, and in fields in winter, while adult males defend year -round feeding territories (Butler 1991). SEXUAL BEHAVIOR Mating system and sex ratio. Mostly mono- gamous. Birds: choose new mates each year (Simpson 1984). Pair bond. Elaborate courtship displays; described in detail by Meyerriecks (1960) and Mock (1976,1979,1980). Much variation between individuals and in sequence of displays. Mock (1976) describes pair formation displays as follows. Stretch display (Fig. 2b): unpaired males extend neck, raise bill toward vertical, and erect neck plumes while exhibiting bright soft parts; neck then retracted, accompanied by a moaning goo-goo call (see Sounds: vocalizations). Snap display: erected head, neck, and breast plumes as head is lowered; mandibles are clapped together and legs flexed when the neck is nearly straight. Wing Preen display: bird leans forward, opens wing slightly, runs bill along edge of primaries. Cirde Flights: heron flies laboriously in wide circle above colony with its neck outstretched. Landing call: when heron returns to nest, neck and head plumes erect; utters a series of croaks (see Sounds: vocalizations). Twig Shake: heron grasps a twig on the nest tree and shakes it side -to -side. Crest Raising: erects black occipital plumes. Fluffed Neck display: heron raises head, erects neck feathers, holds bill at or slightly above horizontal. Upright display: neck and bill extended in a straight line about 45° above the horizontal. Arched Neck display: rapid erection of plumes while curving neck so bill points downwards. Forward display (Fig. 2a) heron extends wrists fromits sides, retracts neck on to back, erects all plumes, then stabs forward with the bill, squawks and claps bill. Bill Duels (Fig. 2c): paired male erects plumage, stands tall, and lunges at the face of its mate with wings held away from its body and a dosed bill. Bill Clappering: rapid clicking of bill tips in mid air, directed toward mate. These displays not rigidly organized into predictable sequences (Mock 1976). Three sequences seen most often by Mock (1976) were the Greeting Ceremony, Stick Transfer, and Nest Relief Ceremony. The Greeting Ceremony occurs when a heron joining its mate on the nest gives the Landing Call. The bird on the nest usually responds with a Full Stretch or, less often, an Arched Neck ROBERT W. BUTLER or Fluffed Neck display. The Stick Transfer sequence occurs when the male brings sticks to its mate. She performs the stretch display and takes the sticks. The male then Bill Clappers the female as she places the stick in the nest. During the Nest Relief sequence an arriving heron utters the Landing Call and its mate stands followed by a Full Stretch. Often the pair Bill Clapper, preen, and even sleep before departure takes place. COPULATION Mostly morning and evening because females ate away from nests midday. Few or no displays (Mock 1976); occurs mostly on nest. Male places one foot gently in centre of female's back: Female leans forward, bends ankles slightly, holds wings slightly away from sides. Male grasps female's humeri with toes and lowers himself onto his tarsi, often while flapping his wings. The female moves her rectrices to one side while the male wags his lowered tail over her cloaca. He grasps her head or neck while copulating, then steps off the female. Extra -pair copulations rare (Cottrille and Cottrille 1958, Brandman 1976, Mock 1976, I. Moul pers. comm.). SOCIAL AND INTERSPECIFIC BEHAVIOR Degree of sociality. Forages alone (Kushlan et al. 1985) or in loose flocks throughout year (see Spacing). Roosts alone or in loose flocks, on the ground during day, above ground at night. Hypothesis that colonies serve as information centres for location of food patches is not well supported (Mock et al. 1988). Play. Nestlings and fledglings stab at inanimate objects; adult play is undocumented. Interactions other than predation with members of other species. Utters mobbing calls toward mammalian and avian predators. Mobbed while flying by nesting gulls and chased by nesting Ospreys (Pandion haliaetus). Bald Eagles; Bayer 1979), Turkey Vultures (Cathartes aura; Kushlan 1978), and (rarely) gulls (Bayer 1985b, Quinney et al. 1981) steal large food items. In turn, steals food from gulls (Bayer 1985b) and (rarely) fish from Osprey nests (A. Poole pers. comm.). Catches fish driven near shore by Double- crested Cormorants (RW B): Occasionally feeds with other ciconiiformes (Kushlan 1978) and larids (Bayer 1985b). Its'nests used by Canada Geese (Branta canadensis), House Sparrows (Passer domesticus), and Great -horned Owls (Bubo virginianus; Vermeer 1969, RWB). Recovery of beaver (Castor canadensis) populations in rte. U.S. and in s. Canada has benefited this species by providing additional nesting and foraging areas (e.g., N.Y. State; Andrle and Carroll 1988). The Academy of Natural Sciences of Philadelphia GREAT BLUE HERON Predation. Northwestern Crows (Corvus caurinus) and Common Ravens (C. corax) eat unattended eggs (Butler 1989). Predators of nestlings include eagles (Kelsall and Simpson 1979, Koonz 1980, Forbes 1987, Norman et al. 1989), raccoons (Lopinot 1952, Hjertas 1982), bears (Foss 1980, Parker 1980), Turkey Vultures (Meitner 1951) and Red - tailed Hawks (Buteo jamaicensis;Simpson 1984). Colony -sites abandoned after predators kill adults (Butler 1991) and nestlings (Kelsall and Simpson 1979, Simpson et al. 1987). BREEDING MATE SELECTION Little information. Of 5 marked pairs in 1978, all had new mates the following year (Simpson 1984). Herodias and occidentalis form pairs and produce viable offspring with white and blue plumages (Bent 1926, Powell et al. 1989). PHENOLOGY Pair formation. At some northern latitudes, adults gather in flocks on the ground for several days in spring before entering colony -sites ( "gathering grounds" sensu Mock 1976). Birds do not occupy gathering grounds in Texas and Michigan, and some roost at colony -site year - round (Cottrille and Cottrille 1958, Mock 1976). Function of gathering ground unclear. Little published information on dates of pair formation. Earliest adults return to colony -sites as follows: California, Dec (Brandman 1976); Texas, Jan (Mock 1976); British Columbia, mid -Jan on the coast, late Mar interior sites (Butler et al. 1986, RWB, Cannings et al. 1987); Pennsylvania, mid - Feb (E. Bruckner pers. comm.); Alberta, late Mar (Vermeer 1969). Herons . in some colonies apt'to depart from colony -sites with little provocation at this time;: may be inhibited from displaying by high winds and low temperatures (Palmer 1962). Courtship to egg laying early Jan to mid -Mar in California (Brandman 1976), mid -Feb to early Apr in British Columbia (RWB). Great White Heron (occidentalis group) breed year -round but most begin between Sep and Feb (Powell 1983). First brood per season. Figure 3. First eggs: late Feb in Oregon (Henny and Bethers 1971) and n. California (Pratt 1970, Pratt and Winkler 1985); early Mar in s. California (Brandman 1976); 3rd week Mar in Idaho (Collazo 1981); 1st week Apr in s. British Columbia (Butler 1989); mid-Apr in Nova Scotia (McAloney 1973); late Apr in Alberta (Vermeer 1969). The Birds of North America, No. 25, 1992 Imo Primaries Molt Body Breeding Young E Mgration 7:11 Figure 3. Annual cycle of breeding, molt, and migration of Great Blue Herons In British Columbia. Coastal populations are non-migratory. Thick lines equal peak activity, thin lines off peak. A. Poole, P. Stettenheim, and F. Gill, Editors Second brood per season. Unclear how many late -laid Butches are first or second attempts. Two pairs renested after fledging first brood, 5 pairs after death of nestlings in California (Brandman 1976). Most choose new mates after first attempt fails (Simpson 1984). Insufficient time to raise 2 broods at northern latitudes. NEST SITE Nest as single pairs but mostly in colonies (Fig. 4). Mean distance flown from colony to principal feeding sites 2.3 to 6.5 km (Dowd and Flake 1985b, Thompson 1978, Parris 1979, Butler 1991). Two radio - equipped breeding adults travelled 61.8 and 103.7 km in 20 to 25 h (Parris 1979). Number of nests in British Columbia (Butler 1991), Oregon (Werschkul et al. 1977, Bayer and McMahon 1981), and Maine (Gibbs et al. 1987, Gibbs 1991) positively related to area of nearby foraging habitat. Location of colony sites best explained by distribution of foraging habitats (Butler 1991, Gibbs 1991). Site selection also predator - driven; like most other herons, this species generally selects nest sites difficult for mammalian predators to reach, e.g., islands, trees in swamps, high branches, etc. Recovery of beaver (Castor.canadensis)populations in ne. U.S. and s. Canada may have benefited this species by providing a patchwork of large and small swamps and wet meadows — additional nesting and foraging areas (e.g., N.Y. State; Andrle and Carroll 1988). The American Ornithologists' Union ROBERT W. BUTLER Selection process. Males arrive in colony and settle on nests, where they court females (Brandman 1976, Mock 1976). Nest site fidelity weak; 13 of 14 marked individuals choose different nests the following yea (Simpson 1984), but fidelity to the choice of tree species within colonies can be strong (Kelsall and Simpson 1979). Some colonies have long histories of use: 71 yr in Stanley Park, BC (RWB), 37 yi•in Minnesota (M. Partch pers.comm.). Site characteristics. Nests mostly in trees, up to 30 m or more above ground; usually lowland swamp or upland hardwood forest, islands, forest - bordered lakes and ponds, and riparian wood- lands, including conifers. Where trees not available, nests on ground (usually only on predator -free islands; Taverner 1926, Gill and Mewaldt 1979), in bullrushes, on shrubs (Behle 1958, Vermeer 1969), cacti (Rosenberg et al. 1991), sagebrush (Blus et al. 1980), mangroves (generally islands; Powe111983), duck blinds (Palmer 1962), channel markers (Henny and Kurtz 1978, Blus et al. 1980), and artificial nest platforms (Sandilands 1980). Nests Figure 4. Most Great Blue Herons nest in colonies. Drawing by J. Zickefoose. with other species of herons (Custer et al. 1980), waterbirds, and even hawks, owls, and vultures (Mengel 1965, Ryser 1985, Simpson 1987). Nest density varies from 39 to 750 nests /ha in Oregon colonies (Bayer and McMahon 1981). NEST Sticks gathered from the ground, nearby trees, or unguarded nestsprimarily by males and placed on nest by females ( Cottrille and Cottrille 1958, Palmer 1962, Mock 1976). Nest built in 3 d to 2 wk, a platform of twigs with saucer - shaped interior lined with pine needles, moss, reeds, dry grass, mangrove leaves, or small twigs (Cottrille and Cottrille 1958, Palmer 1962, Pratt 1970). Four nests in British Columbia weighed 2,570, 2,600, 2,800 and 4,820 g (Butler 1989). Material added throughout nesting period. Dimensions. Vary greatly, from flimsy new platforms of sticks just 0.5 m diameter to bulky older structures 0.9 to 1.2 m across (Andrle 1988); about 1 m deep in nests used several times. The Academy of Natural Sciences of Philadelphia 10 GREAT BLUE HERON Microclimate. No information. Maintenance or reuse of nests, alternate nests. Nests often reused for many years. Twigs gathered near nest (Pratt 1970) by male (mostly when eggs are laid and hatched) and placed by female (Brandman 1976). May build new nest if early attempt fails (Pratt 1970). EGGS Shape. Eggs of Great Blue and Great White herons similar (Bent 1926). Oval to long oval, long elliptical or subelliptical (Harrison 1978). Size. Length 50.7 to 76.5 mm; breadth 29.0 to 50.5 mm (Bent 1926). Mean length 61.38 to 65.57 mm, breadth 45.07 to 46.49 mm for 5 subspecies described by Palmer (1962). Similar dimensions (6 subspecies) described by Butler (1989); volume 6.3 to 7.0 cm3, estimated fresh weight 67.7 to 79.9 g. Fresh weight in Nova Scotia: 70.4 g in 1977 (SD = 3.8, range 63.5 -79.5, n = 27) and 71.6 g in 1978 (SD = 4.1, range 61.0 -78.0, n = 34; Quinney and Smith 1979). Miniature eggs rarely laid; one from British Columbia 35 x 45 mm (RWB). Eggshell composition. The average concen- tration of calcium in 8 eggshells from the Columbia River, WA, 37.3% (Rickard and Schuler 1990). The average concentration (ug /g) in the same 8 eggs was 6.7 Zn, 170 Sr, 190 Ba, 48 Fe and 9.0 Cu. Color. Dull pale blue; fades slightly with age. See Harrison (1978). Surface texture. Smooth or slightly rough. Egg laying. Eggs laid mostly at 2 d intervals, occasionally 3 d (Vermeer 1969; Pratt 1970, McAloney 1973). INCUBATION Onset of broodiness and incubation in relation to laying. Incubation by male and female (Pratt 1970, Brandman 1976, Mock 1979) began after first egg was laid in California (Pratt 1970). Incubation patch. No information. Incubation period. About 27 d: Alberta, mean = 26.7 d (range 26-27, n = 6 clutches, Vermeer 1969); California, 25, 28, 29, and 29 d (Pratt 1970); Nova Scotia, mean = 27.1 d (range 25-30 d, n =11 nests; McAloney 1973). Parental behaviour. Bouts long; male on nest an average of 10.4 h each day, females 3.5 h/d (n = 7 nests; Brandman 1976). Females generally incubate at night. Adults average 54 min / h sitting on eggs (Pratt 1970). Eggs rolled by parent an average of once every 2 h (Pratt 1970). Hardiness of eggs. No information. HATCHING Preliminary events and vocalizations. No information. A. Poole, P. Stettenheim, and F. Gill, Editors The Birds of North America, No. 25, 1992 Shell breaking and emergence. Chicks take 48 h or less to become completely free from egg (Quinney 1982). Time between hatching of first and last eggs (when all eggs hatched) ranged from 2 to 6 d (mean = 3 d; McAloney 1973) and 4 to 8 d (Quinney and Smith 1979) in Nova Scotia, 5 to 8 d in California (Pratt 1970, Brandman 1976). Some hatch on same day (Quinney 1982). In Texas, first egg hatches an average 1.6 d before second egg which hatches 1.7 d before third, which in turn hatches 1.9 d before fourth egg (Mock 1978a). Parental assistance and disposal of eggshells. Adults throw most eggshells from nests soon after hatching (Brandman 1976, Bayer 1982, Butler 1989,1991). YOUNG BIRDS Condition at hatching. Semi - altricial; back, head, and sides of Great Blue chicks covered in pale gray down, especially bushy on crown (Harrison 1978). Wings, bill, and legs pinkish gray and mostly unfeathered. Eye bluish and open. Calling (sounds like tik- tik -tik) commences within minutes of leaving the egg. Mean weight at hatching in 2 Nova Scotia colonies: 49.3 g (SD = 3.2, n = 5) and 51.8 g (SD = 7.1, n = 39; Quinney and Smith 1979). Great White chicks covered in white down at hatching. Growth and development. Mass increase. Quinney (1982) weighed and measured growth of chicks up to 30 d of age. Last hatching chicks grew slower than earlier hatching nestmates. Mass increase was nearly linear in first 30 d of age. No measurable sexual dimorphism at least before 28.5 d. Sixteen 45-d -old chicks weighed 86% of adult weight. Asymptote of growth curve of hand - reared chicks was 2.3 kg (D. Bennett pers. comm.). Growth of body parts. Culmen, tarsus, proximal wing length, wing chord and length of tenth primary increase linearly. However, growth over the entire nestling period is probably best explained by logistic growth curve (Quinney 1982). Control of body temperature: Little information. Gular pouch fluttering at 18 d (Pratt 1970) positively correlated with ambient temperature (Dowd and Flake 1985a). Sunbathe at 27 d (Pratt 1970). Behavior and locomotion. Described by Pratt (1970) and Dowd and Flake (1985a). Preening starts at 6 d of age, stagger to their feet at 14 d and walk steadily at 21 d. Brood reduction, prevalent in many nests, proposed to be the outcome of aggression precipitated by size of prey items delivered to chicks by parents (Mock 1985, 1986, 1987). Wing - flapping starts in fourth week and short hops made into branches near nest at 7 wk. Sustained flights begin at 60 d. Recognizes parents approaching colony at 9 wk. Depart nest at mean The American Ornithologists' Union of 81 d (range 64-91 d), chicks of late - nesting pairs at mean of 67 d. PARENTAL CARE Brooding. Shared by both parents (see Behavior self maintenance). Begins immediately after hatching and lasts 3 to 4 wk (RWB). Feeding. Both parents feed chicks (see Behavior. self maintenance). Newly hatched chicks peck at adult's bill, the nest, and each other (Pratt 1970). Adult stands on rim of nest and places food in open bill of chicks ( Cottrille and Cottrille 1958). By end of second week, chicks grasp adult's bill momentarily; by the end of the third week, until they leave nest, chicks pull adult's bill into the nest (Pratt 1970). Sometimes they reach into adult's open mouth (Cottrille and Cottrille 1958). Largest chicks get most food (Mock 1985, 1986, 1987). Chicks also eat regurgitated food dropped by parents in nest for up to about 30 d after hatching, then take food directly from parents (Mock 1987). Food remains eaten by parents. Aggression between nestmates is low and not related to brood size (Mock et al. 1987a). Brood reduction is apparently not mediated by hunger of nestmates but by size of prey items delivered by parents (Mock et al. 1987b). Chicks fed small prey compete to monolopize food boluses and fight with nest mates more frequently than chicks fed large prey items. Two - day -old chicks fed 10 times in 13 h, 6-d- old chicks more variable: 6 times in 13 h, twice in 15 h (Pratt 1970). Number of feeding visits /h peaked at about 29 d after hatch, lowest rate several weeks before fledging (Dowd and Flake 1985b). Nest sanitation. Most eggshells thrown from nests (Brandman 1976). Feces, partly eaten prey, and dead chicks remain in nest and on ground below. Chicks sometimes regurgitate into nest or onto ground below when disturbed (RWB). Parental carrying of young. Not known to occur. COOPERATIVE BREEDING Not known to occur. BROOD PARASITISM Not known to occur. FLEDGLING STAGE Departure from the nest. Age at first flight: Alberta, 52.6 d (range 51 -54 d, Vermeer 1969), 7 to 8 wk in Oregon (Werschkul et a1.1977) and Nova Scotia (Quinney and Smith 1979). First flight as early as 60 d, mean of 81 d, in California (Pratt 1970). • Growth. See above: Breeding: young birds. ROBERT W. BUTLER 11 Associations with parents or other young. Fledglings return to nest to be fed by adults for a further 3 wk ( Quinney and Smith 1979). Fledglings in British Columbia follow adults from the foraging grounds to the nest to be fed, or return to nest alone (RWB). Ability to get around, feed, and care for self. First flights clumsy but improve rapidly. Foraging skills poorer than adults (Quinney and Smith 1980, Butler 1991). Strike and pace rates similar to adults but capture rates about half of adults during 2 mo following nest departure ( Quinney and Smith 1991, Butler 1991). IMMATURE STAGE Independent of adults a few weeks after first flight. Wander in late summer. Forage with adults on beaches in British Columbia until about Oct, after which juveniles feed in grasslands and marshlands (Butler 1991). DEMOGRAPHY AND POPULATIONS MEASURES OF BREEDING ACTIVITY Age at first breeding; intervals between breeding. Most apparently breed in second spring (i.e. > 22 mo), based upon plumage, but this conclusion might reflect ability of observers to age herons. Juvenile plumaged birds have attempted to breed (Pratt 1973, RWB). Clfitchf2 to 6 eggs; generally increases with latitude. Great Blue: California, mean = 3.16, SE = 0.04, n = 297 (Pratt and Winkler 1985); Oregon mean = 4.19, SD not given, n = 32 (Henny and Bethers 1971); Nova Scotia, mean = 4.2, SD = 0.8, n = 36 (McAloney 1973); Quebec, weighted mean = 4.1, n = 77 (data in DesGranges 1979); British Columbia, mean = 4.2, SD = 0.6, n = 23 (Butler 1989); Alberta, mean = 5.0, SD not given, n = 11 (Vermeer 1969). Decreases through season (Pratt and Winkler 1985). Great White: Florida, mean = 2.9, SD = 0.6, n = 64 (Powell 1983); Annual and lifetime reproductive success. Annual success based on number of nests with eggs varies widely. Most authors, however, report only the number of fledged young per successful nest (nest fledging young), which shows small annual variation: mean of means 2.3 young /nest (SD = 0.30, n = 16 studies, RWB). Future studies should determine number of fledglings raised per nesting attempt. Lifetime reproductive success unknown. Most researchers have been unable to see into nests so causes of chick loss remain poorly known. The average reproductive success (mean number The Academy of Natural Sciences of Philadelphia 12 GREAT BLUE HERON of chicks fledged per successful nest /mean dutch size) is 62.5% (n = 8 studies, Florida to Alberta). LIFE SPAN AND SURVIVORSHIP Mortality estimates from band recovery data: first year 69.0 %, second year 36.3%, subsequent years 21.9% (Henny 1972). Varies regionally (Bayer 1981b). Oldest banded heron 23 yr (RWB). Herons banded on refuges in U.S. appeared to live longer than herons banded off refuges (Bayer 1981b). MORTALITY AND DISEASE Diseases and body parasites. Giardia found in adult male in New York (Georgi et al. 1986). Also carries the nematode Eustrongylides ignotus, which can kill herons (Locke 1961). Transmission of fish pathogenic viruses may occur via the Grey Heron (A. cinerea, Peters and Neukirch 1986). RANGE Initial dispersal from natal site. Little information. Band recoveries suggest that few yearlings return to natal areas (Henny 1972). Fidelity to breeding and winter home range. No information, but some adults hold feeding territories through winter Bayer 1978, Butler 1991). Figure 5. Median number of Great Blue Herons per region in Christmas Bird Counts reporting L 1 heron, winter 1991, U.S. and Canada. A. Poole, P. Stettenheim, and F. Gill, Editors The Birds of North America, No. 25, 1992 Dispersal from breeding site or colony. Disperse away from colonies after breeding (Gill and Mewaldt 1979, Pratt 1970, Butler 1991), along coast and up rivers in Texas (Telfair and Swepston 1987). Band recoveries suggest dispersal in all directions 2 to 3 months post - breeding, then southward movement in autumn; one banded juvenile from U.S. found in Belize (Henny 1972). Home range. Little known outside breeding season. Breeding adults range within about 30 km of colony, most much closer (Parris 1979, Butler 1991). POPULATION STATUS Estimates or counts of density. Coastal breeding populations of Great Blue Heron on U.S. East Coast, Gulf Coast, and Great Lakes show highest densities in Texas, Mississippi delta, w. Florida, and Chesapeake Bay (Spendelow and Patton 1988). Stable or increasing in Tennessee (since 1974; Fleming et al. 1984) and Illinois (since 1982; Kleen 1987), probably stable on coast of British Columbia (since 1982; Butler 1989) and n. Washington (Murphy 1988), but many colonies abandoned after a :few years. Increasing in Michigan (since early .1980s; W. C. Scharf, unpubl. data) and probably in Louisiana (5 censuses over 37 yr; Ogden 1978). In New York, increased 1952 to 1962 (Bull 1964) and since 1962 (Andrle 1988). Decreased after dams built on Colorado River (Rosenberg et al. 1991), probably declined in Iowa (Dinsmore et a1.1984) and in Illinois (Robbins et al. 1986). Great White Heron have rebounded since the 1930s and may be stable or still increasing (Ogden 1978).Figure 5 shows winter density. Population numbers. About 32% of all breeding Great Blue Herons (36,248 birds) along coast from Texas to Maine and in U.S. Great Lakes were on Atlantic Coast, < 10% in Florida, 40% in Louisiana and Texas, and 18% along the Great Lakes (Spendelow and Patton 1988). About 12,211 herodias nests in 248 colonies and 34 isolates in Ontario in early 1980s (Peck and James 1983), minimum of 13,022 pairs in Ontario in 1980 to 1981 (Dunn et al. 1985). On Great Lakes in Michigan, 1,064 nests tallied in 32 colonies in 1987 (Scharf, unpubl. data). About 5,340 nesting pairs in 35 colonies in Illinois (IQeen 1987). Between 1967 to 1972, 4,000 nests counted in 56 colonies in Canadian Prairies (Vermeer 1973). Little information elsewhere in heron's range. About 1,500 individual Great White Herons estimated for Florida Bay in 1985 (Powell et a1.1989). The American Ornithologists' Union POPULATION REGULATION Severe winters might reduce northern popula- tions when foraging sites freeze (Blus and Henny 1981). Populations in Florida decimated by hurricanes, but recovered to former levels (Powell et al. 1989). Evidence for regulation during the breeding season weak or non - existent. Nest sites not limiting in general; number of breeding pairs in colonies positively related to the area of nearby wetlands (Bayer and McMahon 1981, Gibbs et al. 1987), suggesting food supply a critical limitation, perhaps not surprising in a species so strongly territorial. CONSERVATION AND MANAGEMENT EFFECTS OF HUMAN ACTIVITY Contaminants. Eggshell thickness correlated negatively with DDE (Blus et al. 1980, Laporte 1982, Fleming et al. 1984) but not PCB (Vermeer and Reynolds 1972, Ohlendorf et al. 1979, Fleming et al. 1984, but see Blus et al. 1980), although PCB levels were high in a colony that failed in Quebec (Laporte 1982). Mean eggshell thickness has declined compared to pre-1947 levels (Anderson and Hickey 1972): 5.3% to 13.1% (12 colonies, Laporte 1982; 2 colonies, Blus et al. 1980;13 colonies, Bayer 1982; 4 colonies, Fleming et al. 1984). Eggshells broken during incubation an average of 14% to 17% thinner than pre-1947 levels (Faber et al. 1972, Bayer 1982). Wide variation in intraclutch eggshell thickness (Fleming et al. 1984). No clear evidence of adverse effects on reproductive success (Blus et al. 1980, Fleming et al. 1984), but measures of success often too crude to detect subtle differences (see Demography and Populations: measures of breeding activity). Induction of liver enzymes (Bellward et al. 1990), reduced growth and development of young (Hart et a1.1991), and possible reproductive failure (Elliott et al. 1988, 1989) associated with colonies highly contaminated with dioxins (but see Moul 1990). High con- centrations of dieldrin (Ohlendorf et al. 1981) and endrin (Ohlendorf et al. 1979) suspected to be lethal to Great Blue Herons. Disturbance. Nest and colony abandonments increase with increased visits by humans (Drapeau et al. 1984) and with road building and logging activity within 0.5 km (Werschkul et al. 1976). Some colonies splinter and attempt to settle nearby following abandonment (Parker 1980). Response to disturbance can vary between sites and time of breeding season (Vos et a1.1985). Early in season, herons flush easily from nests with slightest disturbance; after eggs, they fly reluctantly and ROBERT W. BUTLER 13 Figure 6. Great Blue Heron at rest. Drawing by AIIan Brooks; US Fish & Wildlife Service art collection, archives of the Academy of Natural Sciences (Stewart Library), Philadelphia. return quickly to nests; few flush when chicks in nest. Birds habituate to non - threatening repeated activities (Anderson 1978, Parker 1980, Vos et al. 1985). Colonies are probably dynamic in areas of high disturbance with individuals and entire colonies relocating between years. Most studies recommend a minimum 300 m buffer zone from the periphery of colonies in which no human activity should take place during courtship and nesting seasons, with the exception of scientific study (reviewed by Butler 1992); however, the most easily disturbed herons left nests in a colony in British Columbia when Butler (1992) approached on foot within 200 m early in the season. Other colonies were less easily disturbed early in the season. This species might have suffered more from loss of wetlands than from overt human activities (English 1978, Rosenberg et al. 1991) and benefitted from open fish ponds and hatcheries. Populations suffered from shooting in past but have not retracted from former range. Attitudes have changed; conspicuousness of this bird that led to its hunting early in the century may now act The Academy of Natural Sciences of Philadelphia 14 GREAT BLUE HERON in its favour. Protection of colony- sites, feeding grounds, and suitable winter habitats probably most important. Screening protects fish in ponds and hatcheries from this and other herons (Mott 1978). APPEARANCE MOLTS AND PLUMAGES Juvenal plumage. Great Blue Heron: according to Palmer (1962) pushes out down so that chicks are well feathered by one third grown. Most of Basic I plumage of juvenile acquired in autumn and retained for > 1 yr. Basic I plumage. Slaty crown sometimes showing small white basally or totally white feathers, grayish rather than brownish neck, white sides of face, dark neck feathers, and narrow sometimes elongated mantle feathers. Cinnamon colored feather edging on wing coverts (RWB). Alternate I plumage. Acquired in first winter; characterized most notably by molt of new crown feathers. Basic II plumage. Worn by yearlings (13-24 mo or longer); gray or gray - streaked crown (i.e. not all white), blackish sides of crown, and short occipital plumes, elongated feathers on sides of neck and mantle but shorter than in adults, and black shoulder patch. Cinnamon feather edging on wing coverts sometimes present (RWB). Prebasic II molt of all feathers in summer or autumn, completed in autumn. Molt of throat plumes begins after eggs are laid (Apr -May), of flight feathers in Jun in British Columbia (RWB). Alternate II plumage. Second winter composed of at least new forehead, crown feathers, wing covert and some back feathers. Molts probably proceed distally among primaries, proximally in secondaries, and centripetally in rectrices, as in Grey Heron (A. cinerea; Milstein et a1.1970). Great White Heron: wholly white in all stages. Molt sequence probably similar to herodias (Bent 1926, Palmer 1962). Description. Juveniles (0-12 mo) brownish compared to adults; crowns are gray (may have 1 cm'- white patch at 8 mo); no body plumes; rust - brown edging to back and wing coverts; upper maxillary is brownish, lower is yellowish. Yearlings (13-24 mo) resemble adults but have more gray on forehead and crown; rust -brown edges to wing coverts; plumes on breast are short or absent. Adults, Great Blue Heron: sexes similar, but males on average slightly larger than females (Table 1). Adults (> 24 mo) slaty-gray on wings, A. Poole, P. Stettenheim, and F. Gill, Editors The Birds of North America, No. 25, 1992 back and sides of neck; primaries darker than rest of wing; black, white and rusty streaks on front of neck; sides black and rust, black patch near bend of folded wing; underparts streaked with black, white belly and under tail; face and crown white with wide black occipital bands terminating in usually 2 plumes; yellowish bill long and tapered; legs brownish or greenish; plumes extend from sides of neck, breast and over back; middle toe pectinated. 70% of adults in British Columbia can be sexed with 95% confidence using length of exposed culmen (Simpson 1984, Butler et a1.1990). BARE PARTS Breeding season. In Florida, bright orange or red bills, bright lime -green lores, bright red legs and feet and yellow iris (Meyerriecks 1960). In Minnesota and Texas, bills mostly yellow, bright cobalt -blue lores, reddish legs; irides deeper yellow (Mock 1976). British Columbia: bright yellow bills, blue-green lores, and greenish -yellow legs (RWB). Maryland: red bills, blue lores, reddish legs (Dolesh 1984). Color fades soon after egg laying begins. Legs and bills "reddish" in Nova Scotia (Quinney and Smith 1979). Nonbreeding season. Great Blue and Great White in Florida have dull yellow bills, pale grayish -blue lores, and yellow irides; legs and feet yellowish -green in former, brownish or greenish- black in latter (Meyerriecks 1960). In British Columbia, bills dorsally yellowish -brown and ventrally dull yellow, lores pale grayish -blue, legs brownish - green, feet yellow - green, irides yellow (RWB). MEASUREMENTS Adult males slightly larger than adult females (Table 1). Little information on Great White Heron, suggesting similar to Great Blue (Palmer 1962). ACKNOWLEDGEMENTS I thank Darin Bennett, Ed Bruckner, Ian Moul, Max Partch, Bill Scharf and Marilyn Sigman for use of unpublished data, and Range Bayer and Peter Stettenheim for helpful comments on this manuscript. Bob Chandler wrote the section on fossil history, Fred Sheldon the section on systematics. The Christmas Bird Count database depends on input from volunteers across the continent. Cover photo by Arthur Morris. The American Ornithologists' Union ROBERT W. BUTLER 15 Table 1. Length (mm) of exposed culmen and tarsus, and body mass (kg) of Great Blue Herons. Standard deviation and sample size appears in parantheses. Data from British Columbia: Simpson (1984); Nova Scotia and eastern North America: Quinney & Smith (1979); Oregon: Bayer (1981c). LOCATION EXPOSED CL/LMEN TARSUS BODY MASS British Columbia Adult Male Adult Female 137.0 (4.4, 24) 123.9 (4.7, 29) 165.9 (6.7, 24) 152.0 (7.0, 29) 2.48 (0.29, 24) 2.11 (0.34, 29) Nova Scotia Adult Male Adult Female 141.2 (8.6, 9) 136.5 (5.3, 13) 178.7 (11.6, 16) 170.8 (12.2, 15) No data No data Eastern Canada & U.S. No data Adult Males and Females No data 2.23 (0.76, 37) Oregon Adult Juvenile (July) Juvenile (Aug -Dec) Yearling (Jun -Jan) No data No data No data No data No data No data No data No data 2.09 (0, 1) 1.76 (0.29, 4) 1.98 (0.50, 4) 2.22 (0.40, 3) REFERENCES American Birds. 1991.90th Christmas Bird Count. Am. Birds 44: 547 -1042. American Ornithologists' Union. 1983. Check -list of North American birds, 6th ed. Amer. Ornithol. Union, Washington, D.C. Anderson, D. W. and J. J. Hickey. 1972. Eggshell changes in certain North American birds, pp. 514 -540 in Proc. XV Int. Ornithol. Congr., E. J. Brill, Leiden. Anderson, J. M. 1978. Protection and management of wading birds, pp. 99 -104 in Wading birds (A. Sprunt IV, J. C. Ogden, and S. Winkler, Eds.). Natl. Audubon Soc. Res. Rep. No. 7, New York. Andrle, R. F. (Ed.) 1988. The atlas of breeding birds in New York State. Cornell Univ. Press, Ithaca, New York. Bayer, R. D. 1978. Aspects of an Oregon estuarine Great Blue Heron population, pp. 213 -218 in Wading birds (A. Sprunt IV, J. C. Ogden, and S. Winkler, Eds.). Natl. Audubon Soc. Res. Rep. No. 7, New York. Bayer, R. D. 1979. Bald Eagle -Great Blue Heron interactions. Murrelet 60: 31 -33. Bayer, R. D. 1980. Social differences in defecation of Great Blue Herons (Ardea herodias). Auk 97: 900- 901. Bayer, R. D. 1981a. Great Blue Herons "mousing" in western Oregon. Murrelet 62: 91. Bayer, R. D. 1981b. Regional variation of Great Blue Heron longevity. J. Field Ornithol. 52: 210 -213. Bayer, R. D. 1981c. Weights of Great Blue Herons at the Yaquina estuary, Oregon. Murrelet 62: 18-19. Bayer, R. D. 1982. Great Blue Heron eggshell thickness at Oregon estuaries. Wilson Bull. 94: 198 -201. Bayer, R. 1984a. Vocalizations of Great Blue Herons at Yaquina Estuary, Oregon. Colonial Waterbirds 7: 35-44. Bayer, R. 1984b. Foraging ground displays of Great Blue Herons at Yaquina Estuary, Oregon. Colonial Waterbirds 7: 45-54. Bayer, R. D. 1985. Bill length of herons and egrets as an estimator of prey size. Colonial Waterbirds 8:104- 109. Bayer, R. D. 1985a. Shiner perch and Pacific Staghorn sculpins in Yaquina Estaury, Oregon. Northwest Sci. 59: 230 -240. Bayer, R. D. 1985b. Interactions between Great Blue Herons and gulls. Wilson Bull. 97: 538 -541. Bayer, R. D. and E. McMahon. 1981. Colony sizes and hatching synchrony of Great Blue Herons in coastal Oregon. Murrelet 62: 73-79. Becker, J. J. 1982. Birds of the Pleistocene American Falls Local Fauna, Idaho. Condor 84: 449. Becker, J. J. 1984. Additions to the late Pleistocene avifauna of Bradenton, Manatee Couty, Florida. Florida Scientists. 47: 203. Becker, J. J. 1985a. A late Pleistocene (Wisconinan) avifauna from West Palm Beach, Florida. Bull. Brit. Orn. Club. 105: 38. Becker, J. J. 1985b. Fossil Herons (Ayes: Ardeidae) of the late Miocene and early Pliocene of Florida. J. Vert. Paleontol. 5: 25. Becker, J. J. 1986. An early heron (Ayes, Ardeidae, Ardea) from the middle Miocene of Nebraska. J. Paleontol. 60: 968. Behle, W. H. 1958. The bird life of Great Salt Lake. Univ. Utah Press, Salt Lake City. The Academy of Natural Sciences of Philadelphia 16 GREAT BLUE HERON - Bellward, G. D., R. J. Nordstrom, P. E. Whitehead, J. E. Elliott, J. E. Bandiera, S. M. Dworschak, T. Chang, S. Forbes, B. Cadario, L. E. Hart, and K. M. Chen. 1990. Comparison of polychlorinated diben- zodioxin and dibenzofuran levels with hepatic mixed - function oxidase induction in Great Blue Herons. J. Toxicol. Environ. Health 30: 33-52. Bent, A. C. 1926. Life histories of North American marsh birds. U.S. Natl. Mus. Bull. 135. Berger, A. J. 1976. Hawaiian bird life. Univ. Hawaii Press, Honolulu. Black, B. B. and M. W. Collopy. 1983. Nocturnal activity of Great Blue Herons in a north Florida saltmarsh. J. Field Ornithol. 53: 403 -406. Blus, L. J., C. J. Heitny, and T. E. Kaiser. 1980. Pollution ecology of breeding Great Blue Herons in the Columbia Basin, Oregon and Washington. Murrelet 61: 63-71. Blus, L. and C. Henny. 1981. Suspected Great Blue Heron population decline after severe winter in the Columbia Basin. Murrelet 62: 16-18. Bock, W. J. 1956. A generic review of the family Ardeidae (Ayes). Am. Mus. Novitates 1779: 1-49. Brandman, M. 1976. A quantitative analysis of the • annual cycle of behavior in the Great Blue Heron (Ardea herodias). Ph.D. diss., Univ. Calif., Los Angeles. Brock, E. M. 1959. Occurrence of the Great Blue Heron in arctic Alaska. Condor 61: 59-60. Brodkorb, P. 1955. The avifauna of the Bone Valley Formation. Florida Geological Survey. Report of Investigations 14: 17. Brodkorb, P. 1963. Catalogue of fosil birds. Bull. Florida State Mus., Biol. Sci. 7: 284. Bull, J. 1964. Birds of New York area. Harper and Row, New York. Butler, R. W., B. G. Stushnoff, and E. McMackin. 1986. The birds of the Creston Valley and southeastern British Columbia. Can. Wildl. Serv. Occas. Pap. No. 58, Ottawa. Butler, R. W. 1989. Breeding ecology and population trends of the Great Blue Heron (Ardea herodias fannini) in the Strait of Georgia, pp. 112 -117 in The ecology and status of marine and shoreline birds in the Strait of Georgia, British Columbia (K. Vermeer and R. W. Butler, Eds.). Can. Wildl. Serv. Spec. Publ., Ottawa. Butler, R. 1991. Habitat selection and time of breeding in the Great Blue Heron (Ardea herodias). Ph.D. diss., Univ. of Brit. Col., Vancouver. Butler, R. W., A. M. Breault, and T. M. Sullivan. 1990. Measuring animals through a telescope. J. Field Ornithol. 61: 111-114. Butler, R. W. 1992. 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DesGranges, J -L., P. Laporte, and G. Chapdelaine. 1979. First tour of inspection of Quebec heronries, 1977. Can. Wildl. Serv. Progr. Notes No. 93, Ottawa. Dinsmore, J. J., T. H. Kent, D. Koenig, P. C. Petersen, and D. M. Roosa. 1984. Iowa birds. Iowa State Univ. Press, Ames. Dolesh, R. J. 1984. Lord of the shallows -the Great Blue Heron. Natl. Geogr. 165: 540 -554. Dowd, E. M. and L. D. Flake. 1985a. Arrival and departure patterns of Great Blue Herons at a South Dakota colony. Wilson Bull. 97: 374 -378. Dowd, E. M. and L. D. Flake. 1985b. Foraging habitats and movements of nesting Great Blue Herons in a prairie river ecosystem, South Dakota. J. Field Ornithol. 56: 379 -387. Drapeau, P., R. McNeil, and J. Burton. 1984. Influences du derangement humaine et de l'activite du Corrnoran a aigretts Phalacrocorax auritus, sur la reproduction du Grant Heron, Ardea herodias, aux iles de la Madeleine. Can. Field -Nat. 98: 219 -222. Dunn, E. H., D. J. T. Hussell, and J. Siderius. 1985. Status of the Great Blue Heron, Ardea herodias, in Ontario. Can. Field -Nat. 99: 62 -70. Elliott, J. E., R. W. Butler, R. J. Norstrom, and P. E. Whitehead. 1988. Levels of polychlorinated dibenzodioxins and polychlorinated dibenzofurans in eggs of Great Blue Herons ( Ardea herodias) in British Columbia, 1983 -87: possible impacts on reproductive success. Can. Wildl. Serv. Progr. Notes No. 176, Ottawa. The American Ornithologists' Union Elliott, J. E., R. W. Butler, R. J. Norstrom, and P. E. Whitehead. 1989. Environmental contaminants and reproductive success of Great Blue Herons Ardea herodias in British Columbia, 1986 -87. Environ. Pollut. 59:•91 -114. Emslie, S. D. 1992. Two new late Blancan avifaunas from Florida and the extinction of wetland birds in the Plio- Pleistocene, pp. 252 in Papers in avian paleontology, honoring Pierce Brodkorb (K. E. Campbell, Jr., Ed.). Nat. Hist Mus Los Angeles County, Sci. Ser. No. 36. Faber, R. A. 1972. Organochlorine and mercury in Common Egrets and Great Blue Herons. Environ. Pollut. 3: 111 -122. Fleming, W. J., B. P. Pullin, and D. M. Swineford. 1984. Population trends and environmental contaminants in herons in the Tennessee valley, 1980 -1981. Colonial Waterbirds 7: 63-73. Forbes, L. S. 1982. Prey manipulation in the Great Blue Heron. Murrelet 63: 89. Forbes, L. S. 1987. Predation on Great Blue Herons: is it important? Colonial Waterbirds 10: 120 -122. Forbes, L.S. and E. McMackin. 1984. Extreme aggression in Great Blue Herons. Wilson Bull. 96: 318 -319. Forbes, L. S., K. Simpson, J. P. Kelsall, and D. R. Flook. 1985. Reproductive success of Great Blue Herons in British Columbia. Can. J. Zool. 63: 1110 -1113. Foss, E. 1980. A black bear in a Great Blue Heron colony. Murrelet 61: 113. George, C. J. and A. Moore. 1990. Overwintering of the Great Blue Heron at Collins Lake, Schenectady Co., New York. Kingbird 40: 75-82. Georgi, M. R., M. S. Carlisle, and L. E. Smiley. 1986. Giardiasis in a Great Blue Heron (Ardea herodias) in New York State: another potential source of waterborne Giardiasis. Am. J. Epidemiol.123: 916- 917. Gibbs, J. P. 1991. Spatial relationships between nesting colonies and foraging areas of Great Blue Herons. Auk 108: 764 -770. Gibbs, J. P., Woodward, M. L. Hunter, and A. E. Huthchinson. 1987. Determinants of Great Blue Heron colony distribution in coastal Maine. Auk 104: 38-47. Gill, R., Jr. and L. R. Mewaldt. 1979. Dispersal and migratory patterns of San Francisco Bay produced herons, egrets and terns. North Am. Bird Bander 4: 4-13. Godfrey, W. E. 1986. The birds of Canada. Natl. Mus. Canada, Ottawa. Godin, J -G. 1977. A Great Blue Heron preying on shiner perch in deep water. Can. Field -Nat. 91: 88- 90. Gray, P., J. W. Grier, G. D. Hamilton, and D. P. Edwards. 1980. Great Blue Heron colonies in northwestern Ontario. Can. Field -Nat. 94: 182 -184. Guthrie, D. A. 1992. A late Pleistocene avifauna from San Miguel Island, California, pp. 321 in Papers in avian paleontology, honoring Pierce Brodkorb (K. E. Campbell, Jr., Ed.). Nat. Hist Mus Los Angeles County, Sci. Ser. No. 36. Hamblin, N. L. and A. M. Rea. 1985. Isla Cozumel archaeological avifauna, pp. 176 in Prehistoric ROBERT W. BUTLER 17 lowland Maya environment and subsistence economy (M. Pohl, Ed.). Pap. Peabody Mus. Archaeol. Ethnol. Vol. 77. Hancock, J. and H. Elliott. 1978. The herons of the world. Harper and Row, New York. Hancock, J. and J. Kushlan. 1984. The herons handbook. Harper and Row Publ., New York. Harris, M. P. 1973. The Galapagos avifauna. Condor 75: 265 -278. Harrison, C. 1978. A field guide to the nests, eggs and nestlings of North American birds. Collins, Glasgow. Hart, L. E., K. M. Cheng, P. E. Whitehead, R. M. Shat, R. J. Lewis, S. R. Ruschkowski, R. W. Blair, D. C. Bennett, S. M. Bandiera, R. J. Norstrom, and G. D. Bellward. 1991. Dioxin contamination and growth and development in embryos. J. Toxicol. and Environ. Health 32: 331 -344. Heitmeyer, M. E. 1986. Post breeding distribution and habitat use of wading birds in Oklahoma, USA. Colonial Waterbirds 9: 163 -170. Henny, C. J. 1972. An analysis of the population dynamics of selected avian species with special reference to changes during the modern pesticide era. U.S. Fish Wildl. Serv., Wildl. Res. Rep. I. Washington, D.C. Henny, C. J. and M. R. Bethers. 1971. Population ecology of the Great Blue Heron, with special reference to western Oregon. Can. Field -Nat. 85: 205 -209. Henny, C. J. and J. E. Kurtz. 1978. Great Blue Herons respond to nesting habitat loss. Wildl. Soc. Bull. 6: 35-37. Hjertaas, D. G. 1982. Great Blue Herons and raccoons at Nicolle Flats. Blue Jay 40: 36-41. Hoffman, R. D. 1978. The diets of herons and egrets in southwestern Lake Erie, pp. 365 -370 in Wading birds,(A. Sprunt IV, J. C. Ogden, and S. Winkler, Eds.). Natl. Audubon Soc. Res. Rep. No. 7, New York. Hom, C. W. 1983. Foraging ecology of herons in a southern San Francisco Bay saltmarsh. Colonial Waterbirds 6: 37-44. Jensen, J. P. 1932. Great Blue Heron fishing in deep water. Auk 47: 245 -246. Kahl, M. P. 1971. Spread -wing postures and their possible functions in the Ciconiidae. Auk 88: 715- 722. Kelsall, J. P. and K. Simpson. 1980. A three -year study of the Great Blue Heron in southwestern British Columbia. Proc. Colonial Waterbird Grp. 3: 69-74. Kleen, V. 1987. Illinois heron colony surveys, 1987 report. Illinois Birds and Birding 3: 79-82. Koonz, W. H. 1980. Bald Eagle nest in a Manitoba heronry. Blue Jay 38: 47. Krebs, J. R. 1974. Colonial nesting and social feeding as strategies for exploiting food resources in the Great Blue Heron (Ardea herodias). Behaviour 51: 99 -131. Krebs, J. R. and B. Partridge. 1973. Significance of head tilting in the Great Blue Heron. Nature 242: 533- 535. Kushlan, J. A. 1976. Feeding behavior of North American herons. Auk 93: 86-94. The Academy of Natural Sciences of Philadelphia 18 GREAT BLUE HERON Kushlan, J. A. 1978. Feeding ecology of wading birds, pp. 249 -298 in Wading birds (A. Sprunt IV, J. C. Ogden, and S. Winkler, Eds.). Natl. Audubon Soc. Res. Rep. No. 7, New York. Kushlan, J. A., L. G. Morales, and P. C. Frohring. 1985. Foraging niche relations of wading birds in tropical wet savannas, pp. 663 -682 in Neotropical orn- ithology (P. A. Buckle Ed.). Ornithol. Monogr. 36. Laporte, P. 1982. Organochlorine residues and eggshell measurements of Great Blue Heron eggs from Quebec. Colonial Waterbirds 5: 95 -103. Larkins, D. 1989. Heat regulation in the Great - billed Heron Arden sumatrana. Corella 13: 21 -23. Laughlin, S. B. and D.1). Kibbe (Eds.). 1985. The atlas of breeding birds of Vermont. Univ. Press of New England, Hanover. Lish, J. W. 1983. Comparative eye morphology of ciconiiform birds. Ph.D. diss., Univ. Oklahoma, Norman. Locke, L. N. 1961. Heron and egret losses due to verminous peritonitis. Avian Dis. 5: 135 -138. Lopinot, A. C. 1952. Raccoon predation on Great Blue Herons. Auk 68: 235. Mayr, E. and L. L. Short. 1970. Species taxa of North • American birds. Publ. Nuttall Ornithol. Club No. 9: 31. McAloney, K. 1973. The breeding biology of the Great Blue Heron on Tobacco Island, Nova Scotia. Can. Field -Nat. 87: 137 -140. Mehner, J. F. 1951. Turkey Vultures attacking Great Blue Heron. Wilson Bull. 64: 242. Mengel, R. M. 1965. The birds of Kentucky. Ornithol. Monogr. No. 3, Amer. Ornithol. Union. Washington, • D.C. Meyerriecks, A. J. 1960. Comparative breeding behavior of four species of North American herons. Publ. • Nuttal Ornithol. Club, No. 2. Cambridge, Mass. Milstein, P. Le S., L Prestt, and A. A. Bell. 1970. The breeding cycle of the Grey Heron. Ardea 58: 171- 257. Mock, D. W. 1976. Pair formation displays of the Great Blue Heron. Wilson Bull. 88: 185-230. Mock, D. 1978a. Pair - formation displays of the Great Egret. Condor 80: 159-172. Mock, D. 1978b. Siblicide, parent - offspring conflict, and unequal parental investment by egrets and °•herons. Behay. Ecol. Sociobiol. 20: 247 -256. Mock, D. 1979. Display repertoire shifts and "extramarital" courtship in herons. Behaviour 69: 57 -71. Mock, D. 1980. White -dark polymorphism in herons. Proc. 1st Welder Wildl. Found. Symp. 1: 145-161. Mock, D. W. 1985. Siblicidal brood reduction: the prey- , size hypothesis. Am. Nat. 125: 327 -343. Mock, D. 1986. Advantages and disadvantages of egret • ' and heron brood reduction. Evolut. 40: 459-470. Mock, D. W. 1987. Siblicide, parent - offspring conflict, and unequal parental investment by egrets and herons. Behay. Ecol. Sociobiol. 20: 247 -256. Mock, D. W., T. C. Lamey, and D. B. A. Thompson. • 1988. Falsifiability and the information centre hypothesis. Ornis Scand. 19: 231 -248. A. Poole, P. Stettenheim, and F. Gill, Editors The Birds of North America, No. 25, 1992 Montevecchi, W. A. and L. M. Tuck. 1987. New- foundland birds: exploitation, study, and con- servation. Publ. Nuttall. Ornithol. Club, No. 21, Cambridge, MA. Mott, D. F. 1978. Control of wading bird predation at fish - rearing facilities, pp. 131 -134 in Wading birds (A. Sprunt IV, J. C. Ogden, and S. Winkler, Eds.). Natl. Audubon Soc. Res. Rept. No. 7, New York. Moul, I. E. 1990. Environmental contaminants, disturbance and breeding failure at a Great Blue Heron colony on Vancouver Island. M.Sc. thesis, Univ. Brit. Col., Vancouver. Murphy, M. 1988. Status of Great Blue Heron colonies in King County, Washington. Western Birds 19: 37-40 Norman, D. M., A. M. Breault and I. E. Moul. 1989. Bald Eagle incursions and predation at Great Blue Heron colonies. Colonial Waterbirds 12: 143 -230. Ogden, J. C. 1978. Recent population trends of colonial wading birds on the Atlantic and Gulf Coastal plains, pp. 137 -154 in Wading birds (A. Sprunt IV, J. C. Ogden, and S. Winkler, Eds.). Natl. Audubon Soc. Res. Rep. No. 7, New York Ohlendorf, H. M., D. M. Swineford, and L. N. Locke. 1979. Organochlorine poisoning of herons. Proc. Colonial Waterbird Group 3: 176 -185. Ohlendorf, H. M., D. M. Swineford, and L. N. Locke. 1981. Organochlorine residues and - mortality of herons. Pest. Monitor. J. 14: 125 -135. Olson, S. L. 1985. The fossil record of birds, pp. 79 -238 in Avian Biology, Vol. 8, (D. S. Farner, J. R. King, and K. C. Parkes, Eds.). Academic Press, New York. Palmer, R. S. 1962. Handbook of North American birds, Vol. 1. Yale Univ. Press, New Haven, CT. Parker, J. 1980. Great Blue Herons (Ardea herodias) in Northwestern Montana: nesting habitat use and the effects of human disturbance. M.Sc. thesis, Univ. Montana. Parmalee, P. W. 1977. The avifauna from prehistory Arikara sites in South Dakota. Plains Anthropol. 22: 200. Parris, R. W. 1979. Aspects of Great Blue Heron (Ardea herodias) foraging ecology in southwest Lake Erie. M.Sc. thesis, Ohio State Univ., Columbus. Payne, R. B. 1979. Ardeidae, pp. 193 -244 in Checklist of Birds of the World (E. Mayr and G. W. Cottrell, Eds.). Mus. Comp. Zool., Cambridge, MA. Payne, R. B. and C. J. Risley. 1976. Systematics and evolutionary relationships among the herons (Ardeidae). Misc. Publ. Univ. Michigan Mus. Zool. 150: 1-115. Peck, G. K. and R. D. James. 1983. Breeding birds of Ontario. Nidiology and distribution. Vol. 1: non - passerines. Roy. Ont. Mus., Toronto. . Peifer, R. W. 1979. Great Blue Herons foraging for small mammals. Wilson Bull. 91: 63 -631. Peters, F. and M. Neukirch. 1986. Transmission of some fish pathogenic viruses by the heron, Ardea cinerea. J. Fish Dis. 9: 539 -544. Powell, G. V. N. 1983. Food availability and reproduction by Great White Herons Ardea herodias: a food addition study. Colonial Waterbirds 6:139- 147. re z G To rn 8 C 4+ 0. % c N 0 VS a to G A v to .- Body of Water County Location Slow -no wake zones (5 mph maximum speed) in the channels. N Prior to 1975 5 A E 0 . m 0 E W cc -s iv- it a 0 W q c c E w .2 o c co 4 2 T . ffi m 2 • E Z 3 O r E W �� yi6'c co •° m m O E o Q CO a 0 _off W § �g a :60 C 7 E •W �j C_ 7 i N E 9 '�� xE$2 o Wrsr��Y is O W °fa ▪ W S E 3 ° oE2Ea.2 • Ei Ea O i 0 E £ .88 �a R 1 .1, ] w § ' o z°gz°`.°z°z° z°2z, -z° U) Z • W Z .-fV M44)(C �fV Meth W J • m L co aaav�m C' Lochness Lake Golden Lake (2-45) Circle Pines Cooper's Lake (2 -70) Spring Lake Park Slow -no wake zones twenty -four hours a day all year from River Mile 4.4 to 5.2. Ramsey /Andover Z 6 co Y Straight Lake (3-10) Y 0 W CO To Whom It May Concern: January 21, 2002 Summary — The Problem I am very concerned for the unique environment and wildlife in Peltier Lake and I am concerned for the hundreds of users of the Lake. It seems that just two individuals have insisted on constructing and using an illegal slalom water -ski course set in a sheltered part of the lake right next to the Heron rookery where 700 nests have been counted. (It is illegal to leave a course in place overnight without a permit.) I believe their noisy boats and activity disturbed the birds and essentially all the birds left the rookery for the 2001 summer season. In addition, mostly coontail and other beneficial aquatic plants were uprooted by their powerful boats causing environmental damage that can easily be seen from an aerial photo. The lake is only 3 to 4 feet deep in that area and I believe huge amounts of silt were put in suspension by their downward thrusting heavy inboard water -ski boats. The fall of 2001 gave Peltier Lake perhaps the worst water quality I have seen in my 22 years on the Lake and I suspect is could be due to the environmental disregard demonstrated by these two individuals. I have never seen coontail wash ashore like it did in August. After the sheriff finally made them remove the illegal slalom water -ski course, they continued to set up the course during days in the same location. Later they moved into various main parts of the Lake causing hardship and restricting others from full use of Peltier Lake. I saw fisherman and other boats forced to go around the slalom water -ski course they established. They essentially cordoned off their own huge private area in what is supposed to be a public resource. In 1998, these same 2 individuals constructed a slalom water -ski course in the same sensitive place right next to the Heron rookery. At the end of the year bird populations seemed down and there was great concern. A meeting with lakeshore owners and several government agencies was held on April 27, 1999, and an agreement was reached that there would be no water - skiing in the sensitive area near the Heron rookery. On July 7, 2001, I saw these same 2 individuals using a slalom water -ski course again in exactly the same spot. When I approached them with my concern, they said they didn't agree to anything and they could do anything they want. It is crystal clear to me that "informal agreements" do NOT work. Formal protection and regulation is required. Even if these 2 individuals quit their activities, there is always the potential for anyone from anywhere to come in and do the same or worse. All indications are that an entire species, the unique and uncommon Black - crowned Night Heron, has completely vanished since 1998, which exactly coincides with the start of slalom water -ski activity next to the rookery. To me, this is not a coincidence. After foraging at night, Black- crowned Night Herons come back to roost during the daytime and are extremely vulnerable to daytime disturbance which is what would happen with a slalom water -ski course next to the rookery. An entire species being wiped out from the rookery is very disturbing to me. Summary — A Solution I recommend protection with local regulation something like the below be enacted before Marchl5 to include ice -out to ice - covered (otherwise they could setup the day of ice -out). 1. North of the bog poles, all non - powered craft stay at least 500 feet away from the island, and power boats are not permitted. (The mostly beneficial aquatic plants are so thick there that power boats normally have difficulty anyway — it is only 3 to 4 feet deep.) vo craft Per,w«tJea 13 05. Ion -fi t ?vies Poles loo 400t j 0 C Y- 4 f e r 14,11`l'f "e 2. South of the bog poles, fishing boats and other boats going at no -wake speed are permitted to within 100 feet of the island, and water skiing is not permitted within 500 feet of the island. 3. Slalom water -ski courses or any marking off of areas in Peltier Lake is not permitted. (The main part of Peltier Lake is about 2000 feet by 4000 feet and a slalom water -ski course has an impact of over 2000 feet in length with its turn around. Peltier Lake is too small and too unique to justify support of such activity). 4. The island be posted as a bird sanctuary with no trespassing. Signs be added at the boat launch, and buoys, markers, and other signs be located by the island. I hope the various governments pull together and support an action that will help restore and protect the Heron rookery, restore equal access for everyone to the Peltier Lake resource, and preserve and protect the environment from damage. Details — My Background I've lived on Peltier Lake since 1979. In 1989, I started the Peltier Lake Association. Here are some of the accomplishments: • Worked lake level issues when St Paul Water pumped Centerville and Peltier Lake down 6 feet in 1988. • Participated on the Rice Creek Watershed District (RCWD) Citizen Advisory Committee for 3 years and learned much about watersheds. • Began a roughly 4 year Curlyleaf Pondweed harvesting effort for Peltier Lake in 1989. • Motivated RCWD to implement a $90,000 Cleanwater Partnership water quality study for the Centerville and Peltier Lake watershed in 1990. • Worked dam level issues as landowners upstream wanted the dam removed and/or lake levels dropped over winter. • Monitor Peltier Lake water quality for the Metropolitan Council starting in 1989. • Launched an effort to manually dig out recently discovered Eurasian Watermilfoil in Peltier Lake in the fall of 2000. • In 2001, found $5,000 in RCWD grant money to help harvest and attempt to control the exotic Curlyleaf Pondweed in Peltier Lake south of the island. • Participate in the Centerville Park and Recreation Committee and advocate community trails. • Volunteer as an Assistant Scoutmaster. I'm afraid that virtually everything I've worked for in trying to preserve and improve Peltier Lake is in jeopardy Details — Behavior Observed To put the problem into perspective, I believe it is worth describing some observed behavior of the two individuals involved. Both were at the April 1999, meeting where numerous experts explained the sensitivity of the area and to please not ski there. Yet, on July 7, 2001, these same two individuals repeated their previous behavior and set up a slalom water -ski course next to the rookery. When I expressed my concern, they said they didn't agree to anything. Later, when I started asking a few other people on the lake about the situation, I was visited on July 10, 2001, by Jerry Lindner and Cindy Lindner who rudely threatened me with a lawsuit saying I was somehow denying them the right of their disabled child to ski the slalom course. They said the ADA was going to get me. They said they were going to sue the Peltier Lake Association for numerous illegal actions. They each had a sheet of paper and seemed to be reading a script. I said I wanted everything in writing. Then I said you have a ski course, right? He said that it was his ski course and he had it rigged so that it sunk when he wasn't using it so nobody else could use it. It was just for him. I asked how long the ski course had been there. He hesitated and said, I'm not answering that. Then I asked him if it was there right now and he again said, I'm not answering that. On July 23, the Anoka County sheriff said he found the slalom water ski course and told the owner it must be removed. So on July 10, when Jerry Lindner and Cindy Lindner were threatening me and intimidating me with every legal word they could think of, they must have had an illegal water -ski course next to the Heron rookery. Do these actions show respect for regulation or for other people? Regarding the other person, all the years he had his current boat on Peltier Lake, he apparently felt no need to license it. It wasn't until DNR and other vehicles started showing up that he finally got a license for his boat in July. I have seen him drive his boat right next to fishermen when the whole lake was clear (at least now someone has a chance at reading a registration number). I was fishing once myself when he drove by within 50 feet pulling a skier. He happened to come back near us and my daughter called out to him, "Can't you see we're fishing here ?" He said, "There aren't any fish there." I said, "No, not anymore." Do these actions show respect for regulation or for other people? In the spring of 1999, I was canoeing around the island when I discovered a float just under the surface. I pulled it up and saw it was part of the slalom water -ski course from 1998. I looked around and saw another, then another. That's when I stopped looking. I don't know how many more were there, but it seemed like there was already an intention to return. Permits do not extend through the winter. All material must be removed from the lake. In August of 2001, I watched them take down their course in the main part of the lake in the evening. In every case I observed, they only removed the top float. They left all the other apparatus in the water overnight (which is illegal without a permit). They set their course up in the only place where I have ever caught a walleye in the lake. With all kinds of lines, concrete blocks and who knows what down there, I wouldn't go fishing there. And even if they had a slalom water -ski course that sunk when not in use, there would probably be over 1,000 feet of hose, lines and other apparatus under water so you couldn't fish there. My opinion is that strict enforcement of new clear regulations as described before are required to protect the Heron rookery, the environment, and normal users of the lake. Details — The Rookery Peltier Lake contains a large wilderness island that used to be home to Black - crowned Night Herons and a large number of Great Blue Herons, Great Egrets, and other birds. The shoreline areas of the lake to the north, east, and west of this island are also essentially wilderness. These areas are rarely frequented by people because the waters there are only 3 to 4 feet deep with a loose silty mud bottom and the area is packed with mostly beneficial aquatic plants. It is an ideal rookery. On July 7, the day I encountered the slalom water -ski course in use north of the island, I did not see any Herons or Egrets north, east, or south of the island (I could not see west of the island). On July 8, the course was again in heavy use. On July 25, I did a careful survey and could not find any Herons or Egrets anywhere on Peltier Lake west, south, or east of the island. The Herons and Egrets seemed essentially gone for the season. There may be some here and there but for 22 years previous to 1998, I have always seen Great Blue Herons, Great Egrets, and Black - crowned Night Herons from my house from spring to fall essentially every day. They land on the dock or fly to the delta at Clearwater creek about 'h mile away from my house. From my house since July 7, I did not seen any Black- crowed Night Herons, Great Egrets, or Great Blue Herons for at least 3 weeks. Most significant, however, the Black - crowned Night Herons disappeared 3 years ago, exactly corresponding to the establishment of the first slalom water ski course. The summer of 2000 also saw abandonment of the rookery sometime in June. There was speculation that it might have been the I35W road reconstruction, however, it is also worth considering the possibility that there was slalom water -ski activity north of the island. This area is not easy to see from the main part of the lake (which is why the birds like it so well). North, east and west of the island, most vegetation consists of native beneficial aquatic plants such as coontail. An image showing the course cut through these aquatic plants is available to anyone via www.mapquest.com. Type in "Peltier Lake" with zip code 55038, pick one of the results and recenter upward (northward) to the top of the island. Click on the "Aerial Photo" tab. Right click it and do a "save as" and lighten the image to better see the track. The ski track is almost horizontal just north of the island angled slightly upward (see dashed line in figure on page 1) and the right hand end has a counter clockwise curl where the ski boats turn around. Details — Slalom Water -ski Course Impact Almost everyone on the lake lives south of the island with the western shore belonging to Anoka County Parks. The area south of the island is narrow and small. Refer to the map of Peltier Lake with a slalom water ski course drawn to scale. In the center, is the course (with its 22 to 28 buoys) and two turnaround ends. Surrounding it all is a buffer of 150 feet. The total estimated impact footprint for a course with a short turn around is about 2,320 feet by 448 feet and for a course with a long turn around, 2,580 feet by 448 feet (course data from www.iwsf.com). The surface area depicted is for the smaller course and has a impact size of about 25 acres. Estimated Slalom Water Ski Course Impacts On Peltier Lake Slalom water ski course size to scale 150' buffer 2 tum arounds course I Smallest version of official slalom water ski course 2,320' x 449' (largest is about 2,580' x 8 1000 Feet The geography of Peltier Lake does not allow for easy placement of a course without severely hampering other users of the lake. People tend to fish in a trolling circle around the lake south of the island right through the areas where the two vertical courses are shown. With a slalom water -ski course on either the east or west sides of the lake, the fishing area is essentially reduced by almost half. Because Peltier Lake is long and narrow instead of circular, the placement of a slalom water ski course anywhere dramatically affects the ability of others to use the lake. There is no reason tiny Peltier Lake should dedicate a third of its usable surface area to such a resource hogging specialized activity. There are alternatives for those who want a slalom water -ski course. Just across the road, on adjacent Centerville Lake, there is a water ski club that performs at least weekly. They already regularly obtain a permit for a water ski jump and it seems likely that something could be set up with this club to provide a slalom water -ski course in the same rough area. Centerville Lake is much bigger in its usable area. As another alternative, Goose Lake, just south of White Bear Lake, has a permanent slalom water -ski course installation. As boating pressures continue to increase in the future, it will be even more important to be careful that everyone has equal access to lake resources. Not permitting slalom water -ski courses just ensures that everyone can fish, boat, and water ski with equal access to the Peltier Lake resource. It also helps ensure protection of the rookery. Conclusion I'd like to see Peltier Lake given back to Black - crowned Night Herons, the Great Egrets, and Great Blue Herons. I'd like to see that fishermen, simple water skiers, sailors, canoeists, and other boaters have equal access to the Peltier Lake resource. The resources of Peltier Lake need formal protection. This is a unique situation with its rookery and relatively small size and requires unique regulation. The impact of a fishing boat or even a single water ski boat with skier is negligible compared with the 25 acre monster foot print of a 2,320 foot by 448 foot slalom water -ski course. See again the four points listed under "A Soulution" above. Thank you. Wayne LeBlanc Appe Aerial photo taken July 23, 2001 north of island. Swirl in lower center is turn around. July 8, 2001 north of island. August 4, 2001. Slalom course north of island. Setting up course north of island, August 4, 2001. August, 2001. Course in main part of lake (south of the island). Zan ILA _014_ 13_4.4____ B H4_ B B....±. B A E o / \D it t t C ' t r is F ro- 1 1 `� ( / • t ,► .tal t t E I G; I 1 / \ 1 % Q \ E + Ei ki +f 't, ! r f i E I *• / • • • , • • t 4 ° • • F L. , %� \ t o \ !D — • C \ 1//43 t f d •r • -rt—A - *-w- -- 8— �►,t -- 8 --►� -- 8---- +t-- -B --- -�— B --�w —A Slalom ski course dimensions in meters. A =27, B=41, C= 29.347, D= 47.011, E =1.25, F =11.5, G =1.15 Buoy 6 • • • End Gates • 140 -180rn 55rr (Optional) • 55m Turn around dimensions. Turn Reny • Skier Drop Point Coontail aquatic vegetation washed on shore, August 2001. Rotting coontail, August 20001 Bad water quality, August 2001. John K. VonDeLinde Director May 4, 1999 Anoka County Department of Parks & Recreation Wayne LeBlanc Peltier Lake Association 1677 Peltier Lake Drive Centerville, Minnesota 55038 Dear Wayne: 550 Bunker Lake Boulevard NW • Andover, Minnesota 55304 Telephone-2) 757 -3920 • FAX (612) 755 -0230 763 Thank you for setting up the meeting with the Peltier Lake Association April 27. The turnout was exceptional. Everyone's comments were heard and they provided valuable background on the use of the lake and the history behind the development of the Lakeshore. There is obvious passion for the lake and a strong desire to keep a sense of community. Anoka County Parks and Recreation appreciates being a part of the meeting. As everyone is aware, Anoka County is a major landowner along Peltier Lake. We would also be glad to be a part of any future meetings. The St. Paul Waterworks is also a landowner on the lake. Everyone connected to the lake has an interest in issues that apply. This note is meant to provide a review of the decisions made at the April 27 meeting. The consensus was to try the following options: 1. Voluntarily keep any water skiing to the main body of the lake south of the island. There was agreement to avoid skiing in the area north of the island. The request is to keep fast moving boats at least 500 feet from the island. 2. Anoka County Parks and Recreation will work with the water skiing sub- committee to educate the general public in regards to the valuable natural resource the northern third of the lake provides and the importance of limiting any fast- moving water activity in that area. Initially, signs will be placed at the public access to inform and educate. Signs will also be placed on the island shoreline declaring the area as a bird sanctuary. 3. Mayor Kim Sullivan, of Lino Lakes, will approach the sea plane operation in order to educate the business about the heron rookery and the need to avoid the area from a safety perspective for both the planes and the birds. Affirmative Action / Equal Opportunity Employer 4. The Department of Natural Resources will monitor the rookery and the behavior of the variety of nesting birds. Lakeshore owners will also monitor lake use and rookery activity. Perhaps this fall or winter all parties can reconvene to discuss their findings. The residents gathered that evening seem very genuine about working together to allow everyone the opportunity to enjoy the beauty of the natural resources in and around Peltier Lake. Hopefully, they have a better understanding of the rookery and the importance of preserving the habitat needed to support the wildlife. Thank you again for organizing this meeting and for guiding the discussion. Please feel free to contact us with any further information about the water ski sub - committee. Let's hope for a wonderful summer season. Sincerely, Jon Oyanagi Park Operations Manager cc: Kim Sullivan, Mayor, City of Lino Lakes Jim March, City Administrator, City of Centerville Bruce Andersohn, Anoka County Sheriff Jeff Perry, Natural Resource Specialist John VonDelinde, Director of Parks and Recreation A/ctv,te- C_e,3)anc- C7.4' u2.24-tA Gttat•Z- Pre ig ...LLUA L) .1)4'h) V 5"7-.A-• & ,#9 /g6CZLd a(AA.4. Y,A t 4 eionein- M FP-r7CAOF'- Tero&T (// Ver/1€((v\ To 11 6e)- (Zn (w.) )) __ ruce. fienver5ohVt C ' 1) ((1 471-017-'91 "CC-t,a� 142.6 -- 0) 6? ,2?S-, F37 611 --24.) -2 54 • 65-1- earl- ZM' gc ,' Vd ?- -off ()5) 65145 GS-1 -4.o (��� t9 Y S� `-fp6 c I.5) ‘.‘7 9 t‘?i 6c( 6 C3- 43P.\ -sl *7—----- -6a 60-7 qz4 -2o�z. 6.-1 1.1Z? — 2-7 6'57 653- -Mr 4.5-1-42g 6 5/ 5-3 Re Yi Tl-g2- -537.5 Agenda Item 6B Environmental Board Meeting Date: January 30, 2002 Topic: Solid Waste /Recycling Report/July 1, 2001 To December 31, 2001 Summary: The City is required to submit two annual reports each year to Anoka County Integrated Waste. Please find attached summary tables of tonnage and costs submitted to Anoka County for the second half of 2001. The City receives SCORE Grant dollars based on the costs and tonnage reported. Attachments: 1. Recycling Tonnage Report r C 0 1- c � N O Co CO '-- 0, as CC O co (0 t0 L TO CO N m M N N N et 10 M ° c 0 0 OMNI 0) N N O) M - O A co O O O t0 CO 0) O BFI of Minnesota 0 a) N e U a M Cr) O O O O 0) a) Co c co al W N n co 42.7 10790 O CO CO M O I0 co O CO N d.0 • 00 M 1- O 1 Recycling Facility O I ! I summary 1_ I— _____1_ _.I Exaense M00 M 10 00 fR $191.40 $510.00 $185.39 10O0 10 M O K) 1p ao d d .N OI10 M I O ? N t- W 10 t- (0 10 M $85.001 $111.90 $146.78 000 01210! TOD ,-(.0.- ! Cl fA $111.901 $425.00 0 01 T P T A $$87.90 $87.90 $111.90 - fA L5253.44 $111_90 $111.90 $111.90 000000 O TD P 00 O O H O O O h M0ID OO O tO 19 WIW O ts. W 1 Totals 10790 3068 47.38 $5,339.211 Cgrs. 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